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crenulate moonwort, dainty moonwort, scalloped moonwort

botryche champètre, Iowa moonwort, prairie moonwort

Pinnae

to 5 pairs, spreading, well separated, distance between 1st and 2d pinnae not or slightly more than between 2d and 3d pairs, basal pinna pair approximately equal in size and cutting to adjacent pair, broadly fan-shaped, undivided to tip, margins mainly crenulate to dentate, proximal pinnae with 1 or more shallow incisions, apex rounded, apical lobe linear to linear-cuneate, well separated from adjacent lobes, venation like ribs of fan, midrib absent.

to 5(–9) pairs, spreading, usually remote, separated 1–3 times pinna width, in some populations irregularly and extensively fused with considerable webbing along rachis, distance between 1st and 2d pinnae not or slightly more than between 2d and 3d pairs, basal pinna pair approximately equal in size and cutting to the adjacent pair, mostly linear to linear-spatulate, undivided to tip, margins crenulate to dentate, usually notched or cleft into 2 or several segments, apex rounded to acute, venation like ribs of fan, midrib absent.

Trophophore

stalk 0.5–7 mm;

blade yellow-green, oblong, 1-pinnate, to 6 × 2 cm, thin, herbaceous.

stalk usually absent but sometimes broadly tapered to 10 mm in forms with coalesced proximal pinnae;

blade glaucescent, oblong, longitudinally folded when alive, 1-pinnate, to 4 × 1.3 cm, very fleshy.

Sporophores

1–2-pinnate, 1.3–3 times length of trophophore.

1(–2, rarely)-pinnate, 1–1.5 times length of trophophore.

2n

=90.

=90.

Botrychium crenulatum

Botrychium campestre

Phenology Leaves appearing in mid to late spring, dying in late summer; in extremely dry years of shorter duration or not appearing at all. Leaves appearing in early spring and dying in late spring and early summer, long before those of associated moonworts.
Habitat Local in marshy and springy areas Extremely inconspicuous in prairies, dunes, grassy railroad sidings, and fields over limestone
Elevation 1200–2500 m (3900–8200 ft) 50–1200 m (200–3900 ft)
Distribution
from FNA
AZ; CA; ID; MT; NV; OR; UT; WA; WY
[WildflowerSearch map]
[BONAP county map]
from FNA
CO; IA; MI; MN; MT; ND; NE; NY; SD; WI; WY; AB; ON; SK
[BONAP county map]
Discussion

Botrychium crenulatum is commonly associated with B. simplex in California. In the Wallowa Mountains of Oregon it occurs with B. ascendens, B. lunaria, and B. minganense.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Botrychium campestre is one of four moonwort species that commonly produce dense clusters of minute, spheric gemmae at the root bases. Peculiar forms of B. campestre with coalescent pinnae are found on dunes in the vicinity of Lake Michigan.

Of conservation concern.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 2. FNA vol. 2.
Parent taxa Ophioglossaceae > Botrychium > subg. Botrychium Ophioglossaceae > Botrychium > subg. Botrychium
Sibling taxa
B. acuminatum, B. ascendens, B. biternatum, B. boreale, B. campestre, B. dissectum, B. echo, B. gallicomontanum, B. hesperium, B. jenmanii, B. lanceolatum, B. lunaria, B. lunarioides, B. matricariifolium, B. minganense, B. montanum, B. mormo, B. multifidum, B. oneidense, B. pallidum, B. paradoxum, B. pedunculosum, B. pinnatum, B. pseudopinnatum, B. pumicola, B. rugulosum, B. simplex, B. spathulatum, B. virginianum
B. acuminatum, B. ascendens, B. biternatum, B. boreale, B. crenulatum, B. dissectum, B. echo, B. gallicomontanum, B. hesperium, B. jenmanii, B. lanceolatum, B. lunaria, B. lunarioides, B. matricariifolium, B. minganense, B. montanum, B. mormo, B. multifidum, B. oneidense, B. pallidum, B. paradoxum, B. pedunculosum, B. pinnatum, B. pseudopinnatum, B. pumicola, B. rugulosum, B. simplex, B. spathulatum, B. virginianum
Name authority W. H. Wagner: Amer. Fern J. 71: 21. (1981) W. H. Wagner & Farrar: Amer. Fern J. 76: 39, figs. 2, 4, 5. (1986)
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