Botrychium crenulatum |
Botrychium ascendens |
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crenulate moonwort, dainty moonwort, scalloped moonwort |
trianglelobe moonwort, upswept moonwort |
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Pinnae | to 5 pairs, spreading, well separated, distance between 1st and 2d pinnae not or slightly more than between 2d and 3d pairs, basal pinna pair approximately equal in size and cutting to adjacent pair, broadly fan-shaped, undivided to tip, margins mainly crenulate to dentate, proximal pinnae with 1 or more shallow incisions, apex rounded, apical lobe linear to linear-cuneate, well separated from adjacent lobes, venation like ribs of fan, midrib absent. |
to 5 pairs, strongly ascending, well separated, distance between 1st and 2d pinnae not or slightly more than between 2d and 3d pairs, basal pinna pair approximately equal in size and cutting to adjacent pair, obliquely narrowly cuneate, undivided to tip, margins sharply denticulate and often shallowly incised, apex rounded, venation like ribs of fan, midrib absent. |
Trophophore | stalk 0.5–7 mm; blade yellow-green, oblong, 1-pinnate, to 6 × 2 cm, thin, herbaceous. |
stalk 3–10 mm, 1/6 length of trophophore rachis; blade yellow-green, oblong to oblong-lanceolate, 1-pinnate, to 6 × 1.5 cm, thin but firm. |
Sporophores | 1–2-pinnate, 1.3–3 times length of trophophore. |
2-pinnate at base of sporangial cluster, 1.3–2 times length of trophophore. |
2n | =90. |
=180. |
Botrychium crenulatum |
Botrychium ascendens |
|
Phenology | Leaves appearing in mid to late spring, dying in late summer; in extremely dry years of shorter duration or not appearing at all. | Leaves appearing in late spring to midsummer. |
Habitat | Local in marshy and springy areas | In grassy fields, widely scattered |
Elevation | 1200–2500 m (3900–8200 ft) | 0–2500 m (0–8200 ft) |
Distribution |
AZ; CA; ID; MT; NV; OR; UT; WA; WY
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AK; CA; MT; NV; OR; WY; BC; ON; YT
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Discussion | Botrychium crenulatum is commonly associated with B. simplex in California. In the Wallowa Mountains of Oregon it occurs with B. ascendens, B. lunaria, and B. minganense. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Botrychium ascendens is a distinctive little moonwort that grows with B. crenulatum, B. lunaria, and B. minganense. This species and B. pedunculosum are the only grapeferns that often have extra sporangia on the proximal pinnae. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 2. | FNA vol. 2. |
Parent taxa | Ophioglossaceae > Botrychium > subg. Botrychium | Ophioglossaceae > Botrychium > subg. Botrychium |
Sibling taxa | ||
Name authority | W. H. Wagner: Amer. Fern J. 71: 21. (1981) | W. H. Wagner: Amer. Fern J. 76: 36, figs. 1, 2. (1986) |
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