Botrychium biternatum |
Botrychium spathulatum |
|
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southern grapefern, sparse-lobed grapefern, sparselobe grapefern |
botryche à segments spatulés, spathulate botrychium, spatulate moonwort, spoon-shape moonwort |
|
Pinnae | to 7 pairs, usually remote, horizontal, distance between 1st and 2d pinnae not or slightly more than between 2d and 3d pairs, undivided except in proximal 2/3–1/2. |
to 8 pairs, ascending, remote, distance between 1st and 2d pinnae not or slightly more than between 2d and 3d pairs, basal pinna pair approximately equal in size and cutting to adjacent pair, mostly narrowly spatulate to linear-spatulate and rounded or ± 2-cleft, lobed to unlobed to tip, margins mainly entire or occasionally irregularly and shallowly incised, apex rounded-notched, venation like ribs of fan, midrib absent. |
Trophophore | stalk 4–20 cm, 2–2.5 times length of trophophore rachis; blade green to dark green, plane, 2–3-pinnate, to 18 × 28 cm, herbaceous. |
stalk 0–1 mm; blade shiny yellow-green, narrowly deltate, flat, 1-pinnate, to 8 × 2.5 cm, thick, leathery. |
Pinnules | elongate, obliquely lanceolate to narrowly lanceolate, margins nearly parallel and finely denticulate, apex short-acuminate, venation pinnate. |
|
Sporophores | 1–2-pinnate, 2–3 times length of trophophore. |
1–2-pinnate, 1.2–2 times length of trophophore. |
2n | =90. |
=180. |
Botrychium biternatum |
Botrychium spathulatum |
|
Phenology | Leaves green over winter, sporophores seasonal, new leaves appearing in late spring–early summer. | Leaves appearing late spring through summer. |
Habitat | Frequent in low woods and brushy fields | Sand dunes, old fields, and grassy railroad sidings |
Elevation | 0–600 m (0–2000 ft) | 0–2000 m (0–6600 ft) |
Distribution |
AL; AR; DE; FL; GA; IL; IN; KY; LA; MD; MO; MS; NC; NJ; OH; OK; PA; SC; TN; TX; VA; WV
|
AK; MI; MT; AB; BC; NB; NT; ON; PE; QC; YT |
Discussion | Botrychium biternatum often grows with B. dissectum and B. jenmanii. The name B. biternatum was misapplied by L.Underwood to B. lunarioides (W.H. Wagner Jr. 1961). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Botrychium spathulatum has long been confused with the more common B. minganense, with which it often grows in the Lake Superior region. The leaves appear later in B. spathulatum than in B. minganense. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 2. | FNA vol. 2. |
Parent taxa | Ophioglossaceae > Botrychium > subg. Sceptridium > sect. Sceptridium | Ophioglossaceae > Botrychium > subg. Botrychium |
Sibling taxa | ||
Synonyms | Osmunda biternata, B. dissectum var. tenuifolium, B. tenuifolium | |
Name authority | (Savigny) L. Underwood: Bot. Gaz. 22: 407. (1896) | W. H. Wagner: Amer. Fern J. 80: 77. (1990) |
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