Botrychium ascendens |
Botrychium spathulatum |
|
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trianglelobe moonwort, upswept moonwort |
botryche à segments spatulés, spathulate botrychium, spatulate moonwort, spoon-shape moonwort |
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Pinnae | to 5 pairs, strongly ascending, well separated, distance between 1st and 2d pinnae not or slightly more than between 2d and 3d pairs, basal pinna pair approximately equal in size and cutting to adjacent pair, obliquely narrowly cuneate, undivided to tip, margins sharply denticulate and often shallowly incised, apex rounded, venation like ribs of fan, midrib absent. |
to 8 pairs, ascending, remote, distance between 1st and 2d pinnae not or slightly more than between 2d and 3d pairs, basal pinna pair approximately equal in size and cutting to adjacent pair, mostly narrowly spatulate to linear-spatulate and rounded or ± 2-cleft, lobed to unlobed to tip, margins mainly entire or occasionally irregularly and shallowly incised, apex rounded-notched, venation like ribs of fan, midrib absent. |
Trophophore | stalk 3–10 mm, 1/6 length of trophophore rachis; blade yellow-green, oblong to oblong-lanceolate, 1-pinnate, to 6 × 1.5 cm, thin but firm. |
stalk 0–1 mm; blade shiny yellow-green, narrowly deltate, flat, 1-pinnate, to 8 × 2.5 cm, thick, leathery. |
Sporophores | 2-pinnate at base of sporangial cluster, 1.3–2 times length of trophophore. |
1–2-pinnate, 1.2–2 times length of trophophore. |
2n | =180. |
=180. |
Botrychium ascendens |
Botrychium spathulatum |
|
Phenology | Leaves appearing in late spring to midsummer. | Leaves appearing late spring through summer. |
Habitat | In grassy fields, widely scattered | Sand dunes, old fields, and grassy railroad sidings |
Elevation | 0–2500 m (0–8200 ft) | 0–2000 m (0–6600 ft) |
Distribution |
AK; CA; MT; NV; OR; WY; BC; ON; YT
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AK; MI; MT; AB; BC; NB; NT; ON; PE; QC; YT |
Discussion | Botrychium ascendens is a distinctive little moonwort that grows with B. crenulatum, B. lunaria, and B. minganense. This species and B. pedunculosum are the only grapeferns that often have extra sporangia on the proximal pinnae. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Botrychium spathulatum has long been confused with the more common B. minganense, with which it often grows in the Lake Superior region. The leaves appear later in B. spathulatum than in B. minganense. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 2. | FNA vol. 2. |
Parent taxa | Ophioglossaceae > Botrychium > subg. Botrychium | Ophioglossaceae > Botrychium > subg. Botrychium |
Sibling taxa | ||
Name authority | W. H. Wagner: Amer. Fern J. 76: 36, figs. 1, 2. (1986) | W. H. Wagner: Amer. Fern J. 80: 77. (1990) |
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