Flora of North America species comparison

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Botrychium ascendens

Botrychium oneidense

trianglelobe moonwort, upswept moonwort

blunt-lobe grape-fern, blunt-lobed grapefern, botryche d'oneida, botryche du lac onéida
Pinnae

to 5 pairs, strongly ascending, well separated, distance between 1st and 2d pinnae not or slightly more than between 2d and 3d pairs, basal pinna pair approximately equal in size and cutting to adjacent pair, obliquely narrowly cuneate, undivided to tip, margins sharply denticulate and often shallowly incised, apex rounded, venation like ribs of fan, midrib absent.

to 5 pairs, usually remote, horizontal to ascending, distance between 1st and 2d pinnae not or slightly more than between 2d and 3d pairs, undivided except in proximal 2/3–3/4.
Trophophore

stalk 3–10 mm, 1/6 length of trophophore rachis;

blade yellow-green, oblong to oblong-lanceolate, 1-pinnate, to 6 × 1.5 cm, thin but firm.

stalk 2–15 cm, 1.5–2.5 times length of blade rachis;

blade dull bluish green, ± plane, 2–3-pinnate, to 15 × 20 cm, ± leathery.
Sporophores

2-pinnate at base of sporangial cluster, 1.3–2 times length of trophophore.

2–3-pinnate, 1.5–2.5 times length of trophophore.
Pinnules

obliquely ovate, margins finely crenulate to denticulate, apex rounded to acute, venation pinnate.
2n

=180.

=90.

Botrychium ascendens

Botrychium oneidense
Phenology Leaves appearing in late spring to midsummer. Leaves green over winter, sporophores seasonal, new leaves appearing in spring.
Habitat In grassy fields, widely scattered In moist, shady, acidic woods and swamps
Elevation 0–2500 m (0–8200 ft) 0–1200 m (0–3900 ft)
Distribution
from FNA
AK; CA; MT; NV; OR; WY; BC; ON; YT
[WildflowerSearch map]
[BONAP county map]
from FNA
CT; DC; DE; IN; KY; MA; MD; ME; MI; MN; NC; NH; NJ; NY; OH; PA; RI; VA; VT; WI; WV; NB; ON; QC
[WildflowerSearch map]
Discussion

Botrychium ascendens is a distinctive little moonwort that grows with B. crenulatum, B. lunaria, and B. minganense. This species and B. pedunculosum are the only grapeferns that often have extra sporangia on the proximal pinnae.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Botrychium oneidense commonly occurs with B. dissectum and B. multifidum. Young individuals of both may resemble B. oneidense (W.H. Wagner Jr. 1961b).

(Discussion copyrighted by Flora of North America; reprinted with permission.)
Source FNA vol. 2. FNA vol. 2.
Parent taxa Ophioglossaceae > Botrychium > subg. Botrychium Ophioglossaceae > Botrychium > subg. Sceptridium > sect. Sceptridium
Sibling taxa
B. acuminatum, B. biternatum, B. boreale, B. campestre, B. crenulatum, B. dissectum, B. echo, B. gallicomontanum, B. hesperium, B. jenmanii, B. lanceolatum, B. lunaria, B. lunarioides, B. matricariifolium, B. minganense, B. montanum, B. mormo, B. multifidum, B. oneidense, B. pallidum, B. paradoxum, B. pedunculosum, B. pinnatum, B. pseudopinnatum, B. pumicola, B. rugulosum, B. simplex, B. spathulatum, B. virginianum
B. acuminatum, B. ascendens, B. biternatum, B. boreale, B. campestre, B. crenulatum, B. dissectum, B. echo, B. gallicomontanum, B. hesperium, B. jenmanii, B. lanceolatum, B. lunaria, B. lunarioides, B. matricariifolium, B. minganense, B. montanum, B. mormo, B. multifidum, B. pallidum, B. paradoxum, B. pedunculosum, B. pinnatum, B. pseudopinnatum, B. pumicola, B. rugulosum, B. simplex, B. spathulatum, B. virginianum
Synonyms B. ternatum var. oneidense, B. dissectum var. oneidense, B. multifidum var. oneidense
Name authority W. H. Wagner: Amer. Fern J. 76: 36, figs. 1, 2. (1986) (Gilbert) House: Amer. Midl. Naturalist 7: 126. (1905)
Web links 
Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).



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