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trianglelobe moonwort, upswept moonwort

botryche à feuille de matricaire, chamomile grape-fern, daisy-leaf moonwort, daisyleaf grape-fern, matricary grape-fern

Pinnae

to 5 pairs, strongly ascending, well separated, distance between 1st and 2d pinnae not or slightly more than between 2d and 3d pairs, basal pinna pair approximately equal in size and cutting to adjacent pair, obliquely narrowly cuneate, undivided to tip, margins sharply denticulate and often shallowly incised, apex rounded, venation like ribs of fan, midrib absent.

to 7 pairs, somewhat ascending, approximate to slightly remote, distance between 1st and 2d pinnae not or slightly more than between 2d and 3d pairs, basal pinna pair approximately equal in size and cutting to adjacent pairs, spatulate-ovate to narrowly ovate, divided to tip, ultimate segments squarish to linear, rounded to square to acute at apex, margins entire to lobed to fully dissected, apex rounded to acute, venation pinnate.

Trophophore

stalk 3–10 mm, 1/6 length of trophophore rachis;

blade yellow-green, oblong to oblong-lanceolate, 1-pinnate, to 6 × 1.5 cm, thin but firm.

stalk 0–5 mm, to 1/6 length of trophophore rachis;

blade dull, ± glaucescent green, oblong, 1–3-pinnate, to 10 × 9 cm, firm.

Sporophores

2-pinnate at base of sporangial cluster, 1.3–2 times length of trophophore.

1–3-pinnate, 1.3–2.4 times length of trophophore.

2n

=180.

=180.

Botrychium ascendens

Botrychium matricariifolium

Phenology Leaves appearing in late spring to midsummer. Leaves appearing in spring, dying in late summer.
Habitat In grassy fields, widely scattered Old fields, secondary growth woods
Elevation 0–2500 m (0–8200 ft) 0–1200 m (0–3900 ft)
Distribution
from FNA
AK; CA; MT; NV; OR; WY; BC; ON; YT
[WildflowerSearch map]
[BONAP county map]
from FNA
CT; DC; DE; IA; IL; IN; KY; MA; MD; ME; MI; MN; NC; NH; NJ; NY; OH; PA; RI; TN; VA; VT; WI; WV; NB; NF; NS; ON; QC; SPM; Europe
[WildflowerSearch map]
[BONAP county map]
Discussion

Botrychium ascendens is a distinctive little moonwort that grows with B. crenulatum, B. lunaria, and B. minganense. This species and B. pedunculosum are the only grapeferns that often have extra sporangia on the proximal pinnae.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 2. FNA vol. 2.
Parent taxa Ophioglossaceae > Botrychium > subg. Botrychium Ophioglossaceae > Botrychium > subg. Botrychium
Sibling taxa
B. acuminatum, B. biternatum, B. boreale, B. campestre, B. crenulatum, B. dissectum, B. echo, B. gallicomontanum, B. hesperium, B. jenmanii, B. lanceolatum, B. lunaria, B. lunarioides, B. matricariifolium, B. minganense, B. montanum, B. mormo, B. multifidum, B. oneidense, B. pallidum, B. paradoxum, B. pedunculosum, B. pinnatum, B. pseudopinnatum, B. pumicola, B. rugulosum, B. simplex, B. spathulatum, B. virginianum
B. acuminatum, B. ascendens, B. biternatum, B. boreale, B. campestre, B. crenulatum, B. dissectum, B. echo, B. gallicomontanum, B. hesperium, B. jenmanii, B. lanceolatum, B. lunaria, B. lunarioides, B. minganense, B. montanum, B. mormo, B. multifidum, B. oneidense, B. pallidum, B. paradoxum, B. pedunculosum, B. pinnatum, B. pseudopinnatum, B. pumicola, B. rugulosum, B. simplex, B. spathulatum, B. virginianum
Synonyms B. lunaria var. matricariifolium
Name authority W. H. Wagner: Amer. Fern J. 76: 36, figs. 1, 2. (1986) (Döll) A. Braun ex W. D. J. Koch: Syn. Deut. Schweiz. Fl., ed. 2 7: 1009. (1847)
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