Botrychium ascendens |
Botrychium matricariifolium |
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trianglelobe moonwort, upswept moonwort |
botryche à feuille de matricaire, chamomile grape-fern, daisy-leaf moonwort, daisyleaf grape-fern, matricary grape-fern |
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Pinnae | to 5 pairs, strongly ascending, well separated, distance between 1st and 2d pinnae not or slightly more than between 2d and 3d pairs, basal pinna pair approximately equal in size and cutting to adjacent pair, obliquely narrowly cuneate, undivided to tip, margins sharply denticulate and often shallowly incised, apex rounded, venation like ribs of fan, midrib absent. |
to 7 pairs, somewhat ascending, approximate to slightly remote, distance between 1st and 2d pinnae not or slightly more than between 2d and 3d pairs, basal pinna pair approximately equal in size and cutting to adjacent pairs, spatulate-ovate to narrowly ovate, divided to tip, ultimate segments squarish to linear, rounded to square to acute at apex, margins entire to lobed to fully dissected, apex rounded to acute, venation pinnate. |
Trophophore | stalk 3–10 mm, 1/6 length of trophophore rachis; blade yellow-green, oblong to oblong-lanceolate, 1-pinnate, to 6 × 1.5 cm, thin but firm. |
stalk 0–5 mm, to 1/6 length of trophophore rachis; blade dull, ± glaucescent green, oblong, 1–3-pinnate, to 10 × 9 cm, firm. |
Sporophores | 2-pinnate at base of sporangial cluster, 1.3–2 times length of trophophore. |
1–3-pinnate, 1.3–2.4 times length of trophophore. |
2n | =180. |
=180. |
Botrychium ascendens |
Botrychium matricariifolium |
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Phenology | Leaves appearing in late spring to midsummer. | Leaves appearing in spring, dying in late summer. |
Habitat | In grassy fields, widely scattered | Old fields, secondary growth woods |
Elevation | 0–2500 m (0–8200 ft) | 0–1200 m (0–3900 ft) |
Distribution |
AK; CA; MT; NV; OR; WY; BC; ON; YT
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CT; DC; DE; IA; IL; IN; KY; MA; MD; ME; MI; MN; NC; NH; NJ; NY; OH; PA; RI; TN; VA; VT; WI; WV; NB; NF; NS; ON; QC; SPM; Europe
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Discussion | Botrychium ascendens is a distinctive little moonwort that grows with B. crenulatum, B. lunaria, and B. minganense. This species and B. pedunculosum are the only grapeferns that often have extra sporangia on the proximal pinnae. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 2. | FNA vol. 2. |
Parent taxa | Ophioglossaceae > Botrychium > subg. Botrychium | Ophioglossaceae > Botrychium > subg. Botrychium |
Sibling taxa | ||
Synonyms | B. lunaria var. matricariifolium | |
Name authority | W. H. Wagner: Amer. Fern J. 76: 36, figs. 1, 2. (1986) | (Döll) A. Braun ex W. D. J. Koch: Syn. Deut. Schweiz. Fl., ed. 2 7: 1009. (1847) |
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