Botrychium ascendens |
Botrychium lunaria |
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trianglelobe moonwort, upswept moonwort |
botryche lunaire, common moonwort, moonwort grapefern |
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Pinnae | to 5 pairs, strongly ascending, well separated, distance between 1st and 2d pinnae not or slightly more than between 2d and 3d pairs, basal pinna pair approximately equal in size and cutting to adjacent pair, obliquely narrowly cuneate, undivided to tip, margins sharply denticulate and often shallowly incised, apex rounded, venation like ribs of fan, midrib absent. |
to 9 pairs, spreading, mostly overlapping except in shaded forest forms, distance between 1st and 2d pinnae not or slightly more than between 2d and 3d pairs, basal pinna pair approximately equal in size and cutting to adjacent pair, broadly fan-shaped, undivided to tip, margins mainly entire or undulate, rarely dentate, apical lobe usually cuneate to spatulate, notched, approximate to adjacent lobes, apex rounded, venation like ribs of fan, midribs absent. |
Trophophore | stalk 3–10 mm, 1/6 length of trophophore rachis; blade yellow-green, oblong to oblong-lanceolate, 1-pinnate, to 6 × 1.5 cm, thin but firm. |
stalk 0–1 mm; blade dark green, oblong, 1-pinnate, to 10 × 4 cm, thick, fleshy. |
Sporophores | 2-pinnate at base of sporangial cluster, 1.3–2 times length of trophophore. |
1–2-pinnate, 0.8–2 times length of trophophore. |
2n | =180. |
=90. |
Botrychium ascendens |
Botrychium lunaria |
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Phenology | Leaves appearing in late spring to midsummer. | Leaves appearing in spring, dying in latter half of summer. |
Habitat | In grassy fields, widely scattered | Open fields, occasionally forests in southern occurrences |
Elevation | 0–2500 m (0–8200 ft) | 0–3700 m (0–12100 ft) |
Distribution |
AK; CA; MT; NV; OR; WY; BC; ON; YT
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AK; AZ; CA; CO; ID; MA; ME; MI; MN; MT; NH; NM; NV; NY; OR; PA; SD; UT; VT; WA; WI; WY; AB; BC; MB; NB; NF; NS; NT; ON; PE; QC; SK; YT; SPM; Pacific Islands in New Zealand; Greenland; Eurasia; s South America; Australia
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Discussion | Botrychium ascendens is a distinctive little moonwort that grows with B. crenulatum, B. lunaria, and B. minganense. This species and B. pedunculosum are the only grapeferns that often have extra sporangia on the proximal pinnae. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Botrychium lunaria grows with many other species of Botrychium, occasionally hybridizing with them. This species, geographically the most widespread of the moonworts, has notably uniform morphology. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 2. | FNA vol. 2. |
Parent taxa | Ophioglossaceae > Botrychium > subg. Botrychium | Ophioglossaceae > Botrychium > subg. Botrychium |
Sibling taxa | ||
Synonyms | Osmunda lunaria, B. onondagense | |
Name authority | W. H. Wagner: Amer. Fern J. 76: 36, figs. 1, 2. (1986) | (Linnaeus) Swartz: J. Bot. (Schrader) 1800(2): 110. (1801) |
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