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trianglelobe moonwort, upswept moonwort

botryche découpé, cutleaf grapefern, dissected grapefern

Pinnae

to 5 pairs, strongly ascending, well separated, distance between 1st and 2d pinnae not or slightly more than between 2d and 3d pairs, basal pinna pair approximately equal in size and cutting to adjacent pair, obliquely narrowly cuneate, undivided to tip, margins sharply denticulate and often shallowly incised, apex rounded, venation like ribs of fan, midrib absent.

to 10 pairs, approximate to remote, slightly ascending, distance between 1st and 2d pinnae not or slightly more than between 2d and 3d pairs, undivided except in proximal 2/3–3/4.

Trophophore

stalk 3–10 mm, 1/6 length of trophophore rachis;

blade yellow-green, oblong to oblong-lanceolate, 1-pinnate, to 6 × 1.5 cm, thin but firm.

stalk 3–15 cm, 1.5–2.5 times length of trophophore rachis;

blade shiny green, often bronze in winter, plane to convex, 3–4-pinnate, to 20 × 30cm, leathery.

Sporophores

2-pinnate at base of sporangial cluster, 1.3–2 times length of trophophore.

2–3-pinnate, 1.5–2.5 times length of trophophore.

Pinnules

usually obliquely angular–trowel-shaped to widely trowel-shaped to obliquely round-lanceolate to ovate and pointed, margins denticulate to lacerate to coarsely cut halfway or wholly into linear-divergent segments in some populations, venation pinnate.

2n

=180.

=90.

Botrychium ascendens

Botrychium dissectum

Phenology Leaves appearing in late spring to midsummer. Leaves green over winter, new leaves appearing in late spring.
Habitat In grassy fields, widely scattered In variety of habitats, open grassy areas to deep forest
Elevation 0–2500 m (0–8200 ft) 0–1500 m (0–4900 ft)
Distribution
from FNA
AK; CA; MT; NV; OR; WY; BC; ON; YT
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; CT; DC; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MO; MS; NC; NH; NJ; NY; OH; PA; RI; SC; TN; TX; VA; VT; WI; WV; NB; NS; ON; QC; West Indies in the Antilles
Discussion

Botrychium ascendens is a distinctive little moonwort that grows with B. crenulatum, B. lunaria, and B. minganense. This species and B. pedunculosum are the only grapeferns that often have extra sporangia on the proximal pinnae.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Botrychium dissectum is highly variable, even within the same population. In Florida and along the Gulf Coast, the extremely lacerate form is absent, and the blade segments are usually strongly angular, trowel-shaped, and dentate. In eastern Kentucky and central Tennessee in forested valleys, on shale and limestone soils, plants have narrowly linear, somewhat blunt-tipped segments with a more or less whitish gray central line above the veins. This variant, which grows with B. dissectum, may deserve recognition as a distinct species.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 2. FNA vol. 2.
Parent taxa Ophioglossaceae > Botrychium > subg. Botrychium Ophioglossaceae > Botrychium > subg. Sceptridium > sect. Sceptridium
Sibling taxa
B. acuminatum, B. biternatum, B. boreale, B. campestre, B. crenulatum, B. dissectum, B. echo, B. gallicomontanum, B. hesperium, B. jenmanii, B. lanceolatum, B. lunaria, B. lunarioides, B. matricariifolium, B. minganense, B. montanum, B. mormo, B. multifidum, B. oneidense, B. pallidum, B. paradoxum, B. pedunculosum, B. pinnatum, B. pseudopinnatum, B. pumicola, B. rugulosum, B. simplex, B. spathulatum, B. virginianum
B. acuminatum, B. ascendens, B. biternatum, B. boreale, B. campestre, B. crenulatum, B. echo, B. gallicomontanum, B. hesperium, B. jenmanii, B. lanceolatum, B. lunaria, B. lunarioides, B. matricariifolium, B. minganense, B. montanum, B. mormo, B. multifidum, B. oneidense, B. pallidum, B. paradoxum, B. pedunculosum, B. pinnatum, B. pseudopinnatum, B. pumicola, B. rugulosum, B. simplex, B. spathulatum, B. virginianum
Synonyms B. obliquum
Name authority W. H. Wagner: Amer. Fern J. 76: 36, figs. 1, 2. (1986) Sprengel: Anleit. Kenntn. Gew. 3: 172. (1804)
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