Botrychium ascendens |
Botrychium campestre |
|
---|---|---|
trianglelobe moonwort, upswept moonwort |
botryche champètre, Iowa moonwort, prairie moonwort |
|
Pinnae | to 5 pairs, strongly ascending, well separated, distance between 1st and 2d pinnae not or slightly more than between 2d and 3d pairs, basal pinna pair approximately equal in size and cutting to adjacent pair, obliquely narrowly cuneate, undivided to tip, margins sharply denticulate and often shallowly incised, apex rounded, venation like ribs of fan, midrib absent. |
to 5(–9) pairs, spreading, usually remote, separated 1–3 times pinna width, in some populations irregularly and extensively fused with considerable webbing along rachis, distance between 1st and 2d pinnae not or slightly more than between 2d and 3d pairs, basal pinna pair approximately equal in size and cutting to the adjacent pair, mostly linear to linear-spatulate, undivided to tip, margins crenulate to dentate, usually notched or cleft into 2 or several segments, apex rounded to acute, venation like ribs of fan, midrib absent. |
Trophophore | stalk 3–10 mm, 1/6 length of trophophore rachis; blade yellow-green, oblong to oblong-lanceolate, 1-pinnate, to 6 × 1.5 cm, thin but firm. |
stalk usually absent but sometimes broadly tapered to 10 mm in forms with coalesced proximal pinnae; blade glaucescent, oblong, longitudinally folded when alive, 1-pinnate, to 4 × 1.3 cm, very fleshy. |
Sporophores | 2-pinnate at base of sporangial cluster, 1.3–2 times length of trophophore. |
1(–2, rarely)-pinnate, 1–1.5 times length of trophophore. |
2n | =180. |
=90. |
Botrychium ascendens |
Botrychium campestre |
|
Phenology | Leaves appearing in late spring to midsummer. | Leaves appearing in early spring and dying in late spring and early summer, long before those of associated moonworts. |
Habitat | In grassy fields, widely scattered | Extremely inconspicuous in prairies, dunes, grassy railroad sidings, and fields over limestone |
Elevation | 0–2500 m (0–8200 ft) | 50–1200 m (200–3900 ft) |
Distribution |
AK; CA; MT; NV; OR; WY; BC; ON; YT
|
CO; IA; MI; MN; MT; ND; NE; NY; SD; WI; WY; AB; ON; SK |
Discussion | Botrychium ascendens is a distinctive little moonwort that grows with B. crenulatum, B. lunaria, and B. minganense. This species and B. pedunculosum are the only grapeferns that often have extra sporangia on the proximal pinnae. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Botrychium campestre is one of four moonwort species that commonly produce dense clusters of minute, spheric gemmae at the root bases. Peculiar forms of B. campestre with coalescent pinnae are found on dunes in the vicinity of Lake Michigan. Of conservation concern. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 2. | FNA vol. 2. |
Parent taxa | Ophioglossaceae > Botrychium > subg. Botrychium | Ophioglossaceae > Botrychium > subg. Botrychium |
Sibling taxa | ||
Name authority | W. H. Wagner: Amer. Fern J. 76: 36, figs. 1, 2. (1986) | W. H. Wagner & Farrar: Amer. Fern J. 76: 39, figs. 2, 4, 5. (1986) |
Web links |
|