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trianglelobe moonwort, upswept moonwort

southern grapefern, sparse-lobed grapefern, sparselobe grapefern

Pinnae

to 5 pairs, strongly ascending, well separated, distance between 1st and 2d pinnae not or slightly more than between 2d and 3d pairs, basal pinna pair approximately equal in size and cutting to adjacent pair, obliquely narrowly cuneate, undivided to tip, margins sharply denticulate and often shallowly incised, apex rounded, venation like ribs of fan, midrib absent.

to 7 pairs, usually remote, horizontal, distance between 1st and 2d pinnae not or slightly more than between 2d and 3d pairs, undivided except in proximal 2/3–1/2.

Trophophore

stalk 3–10 mm, 1/6 length of trophophore rachis;

blade yellow-green, oblong to oblong-lanceolate, 1-pinnate, to 6 × 1.5 cm, thin but firm.

stalk 4–20 cm, 2–2.5 times length of trophophore rachis;

blade green to dark green, plane, 2–3-pinnate, to 18 × 28 cm, herbaceous.

Sporophores

2-pinnate at base of sporangial cluster, 1.3–2 times length of trophophore.

1–2-pinnate, 2–3 times length of trophophore.

Pinnules

elongate, obliquely lanceolate to narrowly lanceolate, margins nearly parallel and finely denticulate, apex short-acuminate, venation pinnate.

2n

=180.

=90.

Botrychium ascendens

Botrychium biternatum

Phenology Leaves appearing in late spring to midsummer. Leaves green over winter, sporophores seasonal, new leaves appearing in late spring–early summer.
Habitat In grassy fields, widely scattered Frequent in low woods and brushy fields
Elevation 0–2500 m (0–8200 ft) 0–600 m (0–2000 ft)
Distribution
from FNA
AK; CA; MT; NV; OR; WY; BC; ON; YT
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; DE; FL; GA; IL; IN; KY; LA; MD; MO; MS; NC; NJ; OH; OK; PA; SC; TN; TX; VA; WV
[WildflowerSearch map]
Discussion

Botrychium ascendens is a distinctive little moonwort that grows with B. crenulatum, B. lunaria, and B. minganense. This species and B. pedunculosum are the only grapeferns that often have extra sporangia on the proximal pinnae.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Botrychium biternatum often grows with B. dissectum and B. jenmanii. The name B. biternatum was misapplied by L.Underwood to B. lunarioides (W.H. Wagner Jr. 1961).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 2. FNA vol. 2.
Parent taxa Ophioglossaceae > Botrychium > subg. Botrychium Ophioglossaceae > Botrychium > subg. Sceptridium > sect. Sceptridium
Sibling taxa
B. acuminatum, B. biternatum, B. boreale, B. campestre, B. crenulatum, B. dissectum, B. echo, B. gallicomontanum, B. hesperium, B. jenmanii, B. lanceolatum, B. lunaria, B. lunarioides, B. matricariifolium, B. minganense, B. montanum, B. mormo, B. multifidum, B. oneidense, B. pallidum, B. paradoxum, B. pedunculosum, B. pinnatum, B. pseudopinnatum, B. pumicola, B. rugulosum, B. simplex, B. spathulatum, B. virginianum
B. acuminatum, B. ascendens, B. boreale, B. campestre, B. crenulatum, B. dissectum, B. echo, B. gallicomontanum, B. hesperium, B. jenmanii, B. lanceolatum, B. lunaria, B. lunarioides, B. matricariifolium, B. minganense, B. montanum, B. mormo, B. multifidum, B. oneidense, B. pallidum, B. paradoxum, B. pedunculosum, B. pinnatum, B. pseudopinnatum, B. pumicola, B. rugulosum, B. simplex, B. spathulatum, B. virginianum
Synonyms Osmunda biternata, B. dissectum var. tenuifolium, B. tenuifolium
Name authority W. H. Wagner: Amer. Fern J. 76: 36, figs. 1, 2. (1986) (Savigny) L. Underwood: Bot. Gaz. 22: 407. (1896)
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