Botrychium ascendens |
Botrychium biternatum |
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trianglelobe moonwort, upswept moonwort |
southern grapefern, sparse-lobed grapefern, sparselobe grapefern |
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Pinnae | to 5 pairs, strongly ascending, well separated, distance between 1st and 2d pinnae not or slightly more than between 2d and 3d pairs, basal pinna pair approximately equal in size and cutting to adjacent pair, obliquely narrowly cuneate, undivided to tip, margins sharply denticulate and often shallowly incised, apex rounded, venation like ribs of fan, midrib absent. |
to 7 pairs, usually remote, horizontal, distance between 1st and 2d pinnae not or slightly more than between 2d and 3d pairs, undivided except in proximal 2/3–1/2. |
Trophophore | stalk 3–10 mm, 1/6 length of trophophore rachis; blade yellow-green, oblong to oblong-lanceolate, 1-pinnate, to 6 × 1.5 cm, thin but firm. |
stalk 4–20 cm, 2–2.5 times length of trophophore rachis; blade green to dark green, plane, 2–3-pinnate, to 18 × 28 cm, herbaceous. |
Sporophores | 2-pinnate at base of sporangial cluster, 1.3–2 times length of trophophore. |
1–2-pinnate, 2–3 times length of trophophore. |
Pinnules | elongate, obliquely lanceolate to narrowly lanceolate, margins nearly parallel and finely denticulate, apex short-acuminate, venation pinnate. |
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2n | =180. |
=90. |
Botrychium ascendens |
Botrychium biternatum |
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Phenology | Leaves appearing in late spring to midsummer. | Leaves green over winter, sporophores seasonal, new leaves appearing in late spring–early summer. |
Habitat | In grassy fields, widely scattered | Frequent in low woods and brushy fields |
Elevation | 0–2500 m (0–8200 ft) | 0–600 m (0–2000 ft) |
Distribution |
AK; CA; MT; NV; OR; WY; BC; ON; YT
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AL; AR; DE; FL; GA; IL; IN; KY; LA; MD; MO; MS; NC; NJ; OH; OK; PA; SC; TN; TX; VA; WV
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Discussion | Botrychium ascendens is a distinctive little moonwort that grows with B. crenulatum, B. lunaria, and B. minganense. This species and B. pedunculosum are the only grapeferns that often have extra sporangia on the proximal pinnae. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Botrychium biternatum often grows with B. dissectum and B. jenmanii. The name B. biternatum was misapplied by L.Underwood to B. lunarioides (W.H. Wagner Jr. 1961). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 2. | FNA vol. 2. |
Parent taxa | Ophioglossaceae > Botrychium > subg. Botrychium | Ophioglossaceae > Botrychium > subg. Sceptridium > sect. Sceptridium |
Sibling taxa | ||
Synonyms | Osmunda biternata, B. dissectum var. tenuifolium, B. tenuifolium | |
Name authority | W. H. Wagner: Amer. Fern J. 76: 36, figs. 1, 2. (1986) | (Savigny) L. Underwood: Bot. Gaz. 22: 407. (1896) |
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