Bolboschoenus fluviatilis |
Bolboschoenus novae-angliae |
|
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bolboschoenus fluviatile, river bulrush, river tuber-bulrush, scirpe fluviatile |
New England bulrush, New England tuber-bulrush |
|
Culms | 100–200 cm × 5–15 mm. |
to 150 cm × 5–10 mm. |
Leaves | sheaths reaching to middle of culm or higher, fronts convex (to concave) and papery at mouth, veins reaching apex, very rarely diverging leaving triangular, veinless, membranous area; widest blade 7–22 mm wide. |
sheaths reaching beyond or to middle of culm, fronts convex, papery at mouth, veins reaching apex; widest blade 6–13 mm wide. |
Inflorescences | subumbellate, with all or most spikelets solitary or in clusters of 2–3(–8) on 4–12 rays, rays not exceeding 10 cm; involucral bracts that surpass inflorescences 3–6, widest bract 4–15 mm wide. |
subumbellate, all or most spikelets solitary or in clusters of 2–7 on 3–11 rays, rays not exceeding 8 cm; involucral bracts that surpass inflorescences 2–4, widest bract 2–8 mm wide. |
Spikelets | 10–40, ovoid to lanceoloid, 10–25 × 6–10 mm, base cuneate to rounded; scales often loosely imbricate, orange-brown to stramineous, usually obscurely lineolate-spotted, 7–10 × 3–4 mm, membranous and translucent, apex 2-fid 0.5–1 mm deep, awns fairly stout, 2–3 × 0.5 mm wide at base. |
10–40, ovoid or lanceoloid, 15–40 × 5–6(–8) mm; scales tightly appressed or in fruit usually loosely imbricate, medium to dark orange-brown, lineolate-spotted at 15X, 7–9 × 3–4 mm, papery, nearly opaque, apex 2-fid 0.5 mm deep, awn fairly stout, 2–3 × (0.3–)0.5 mm at base. |
Flowers | perianth bristles tightly attached to shed achene, pale brown, very stout, equaling achene; anthers yellow, 4 mm; styles 3-fid. |
perianth bristles weakly attached to shed achene or some to all falling separately, medium brown, 1/2 to equaling length of achene; anthers dark yellow to orange-yellow, 3.5–5 mm; styles (2–)3-fid. |
Achenes | grayish or dark brown, often in patches, obovoid to obpyriform, all nearly equilaterally markedly trigonous to slightly compressed, angles equally rounded, 3.8–5.5 × 2–2.9 mm, apex rounded, beak 0.2–0.8 mm, surface rather dull, exocarp cells usually not evident at 20X; in cross section exocarp much thinner than mesocarp and its cells very small, isodiametric; achene specific gravity greater than water. |
dark to medium brown, color variable often on same achene, shape variable, often in same plant, markedly to obscurely trigonous with equally rounded angles to much compressed-trigonous with abaxial angle broadly rounded, or some biconvex in same spikelet, obovoid, 3–4.3 × 2.3–3.1 mm, apex rounded to nearly truncate, beak 0.1–0.5 mm, surface glossy or glossy with dull patches, entirely clearly cellular at 20X or faintly cellular to noncellular at 30X, often in patches; in cross section exocarp 1/3–2/3 of mesocarp thickness and its cells variably enlarged, 1.5–3 times deeper than wide; achene specific gravity greater to less than water. |
2n | = 94. |
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Bolboschoenus fluviatilis |
Bolboschoenus novae-angliae |
|
Phenology | Fruiting summer. | Fruiting summer. |
Habitat | Fresh shores, inland marshes, coastal estuaries | Slightly brackish coastal shores, estuaries, marshes |
Elevation | 0–2100 m (0–6900 ft) | 0 m (0 ft) |
Distribution |
AL; AZ; CA; CO; CT; DE; IA; ID; IL; IN; KS; MA; MD; ME; MI; MN; MO; MT; ND; NE; NH; NJ; NY; OH; OR; PA; SD; TN; UT; VA; VT; WA; WI; AB; BC; MB; NB; ON; QC; SK; Asia (Japan); Australia; Pacific Islands (New Zealand)
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CT; DE; GA; MA; MD; ME; NJ; NY; RI; VA |
Discussion | Bolboschoenus fluviatilis frequently forms dense, monospecific, often entirely vegetative stands, and it is more common than recorded because vegetative colonies are often overlooked (E. W. Chester and B. E. Wofford 1992). The only record for Alabama is an 1870 collection from the East Fowl River in the Mobile Delta, where the species has not been collected since. It was intentionally introduced into New Hampshire (D. J. Padgett and G. E. Crow 1993). The report from New Mexico by M. L. Fernald (1950) cannot be confirmed because no specimen is known. Putative hybrids with Bolboschoenus maritimus occur in California. Bolboschoenus novae-angliae probably originated from B. fluviatilis × B. robustus (J. Browning et al. 1995). Introgression from B. maritimus and/or B. robustus is suggested by the larger exocarp cells (evident in surface view) in some North American plants. The Eurasian B. yagara (Ohwi) Y. C. Yang & M. Zhan differs from B. fluviatilis in its narrower leaves and smaller achenes. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The only Georgia record is the type of Scirpus maritimus [var.] cylindricus Torrey, which lacks locality data. I have not seen a voucher from Georgia or for the only report from North Carolina (E. O. Beal 1977). In 1913, N. L. Britton and A. Brown treated Bolboschoenus novae-angliae as a species; other authors included it in either Scirpus maritimus or S. robustus until A. E. Schuyler (1975) revived it as a species under the name S. cylindricus (Torrey) Britton [illegitimate, not S. cylindricus (Vahl) Lamarck 1817]. Bolboschoenus novae-angliae is probably of hybrid origin because it is intermediate between B. fluviatilis and B. robustus, especially in its achene structure and perianth bristle persistence (J. Browning et al. 1995), and it is known almost entirely from the zone of sympatry of its putative parents. It is ecologically intermediate between the fresh habitats of B. fluviatilis and the saline habitats of B. maritimus and B. robustus. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23, p. 42. | FNA vol. 23, p. 43. |
Parent taxa | Cyperaceae > Bolboschoenus | Cyperaceae > Bolboschoenus |
Sibling taxa | ||
Synonyms | Scirpus maritimus var. fluviatilis, Scirpus fluviatilis, Schoenoplectus fluviatilis | Scirpus novae-angliae, Schoenoplectus novae-angliae, Scirpus robustus var. novae-angliae |
Name authority | (Torrey) Soják: Cas. Nár. Mus., Odd. Prír. 141: 62. (1972) | (Britton) S. G. Smith: Brittonia 47: 434. (1995) |
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