Boerhavia triquetra |
Boerhavia linearifolia |
|
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creeping sticky stem, slender spiderling |
narrowleaf spiderling |
|
Habit | Herbs, annual; taproot tapered, soft or ± woody. | Herbs, perennial, sometimes slightly woody at base; taproot long, ropelike, ± woody. |
Stems | erect or ascending, occasionally decumbent, moderately or profusely branched, primarily distally, 1–5 dm, minutely puberulent with bent hairs basally, glabrous or minutely puberulent distally. |
usually erect or ascending, occasionally decumbent, sparingly branched throughout, 2–5(–9) dm, hirsute, puberulent, or glandular basally, hair types often mixed, glandular-puberulent, glabrous, or occasionally puberulent and sparsely hispid distally. |
Leaves | mostly in basal 1/2 of plant; larger leaves with petiole 4–8[–15] mm, blade broadly ovate or oval to lanceolate, 10–21[–35] × 6–8[–13] mm (distal leaves smaller, proportionately narrower), adaxial surface usually glabrous, rarely minutely puberulent, abaxial surface paler than adaxial, glabrous, neither surface punctate, base round [obtuse], margins shallowly sinuate, apex acute, obtuse, or round. |
mostly in basal 2/3 of plant; larger leaves with petiole 1–5 mm, blade lanceolate to linear, rarely ovate, 15–35 × 1–15 mm (distal leaves usually longer, narrower), adaxial surface glabrous or sparsely hispid, occasionally glandular-pubescent, sometimes punctate, abaxial surface usually paler than adaxial, glabrous or sparsely hispid, occasionally glandular-pubescent, usually punctate with small patches of large gray to dark brown cells, base obtuse, truncate, or rounded, margins entire and revolute, apex usually acute, sometimes obtuse or rounded. |
Inflorescences | terminal or axillary, forked ca. 3–6 times ± evenly (or clearly unevenly), diffuse, with sticky internodal bands; branches usually terminating in umbels or flowers borne singly, usually well beyond leaves. |
terminal and axillary, forked ca. 2–4 times unequally, open, without sticky internodal bands; branches strongly ascending, terminating in loose, 1–few-flowered, cymose clusters. |
Flowers | pedicel 0.3–4.5 mm; bracts at base of perianth quickly deciduous, 2, linear-lanceolate to lanceolate [ovate], 0.5–0.6 mm; perianth whitish to pale pink or purplish, campanulate beyond constriction, 1 mm; stamens 2–3, included or barely exserted. |
pedicel 1–2(–5) mm; bracts at base of perianth persistent, 3, lanceolate to ovate-acuminate, 1–2 mm; perianth purplish pink, widely funnelform beyond constriction, 4–7 mm; stamens 5(–6), well exserted. |
Fruits | 1–7 per umbel, straw colored or gray-brown, obpyramidal, 2.5–3.1 × 1.3–1.9 mm (l/w: 1.6–2.1), base stipelike, apex round-truncate, glabrous; ribs 3–4(–5), acute or winglike, tapering to pedicel, slightly rugose or undulate near sulci; sulci 0.5–1 times as wide as base of ribs, coarsely transversely rugose, smooth or very faintly papillate. |
borne singly (or 2–3 in open clusters), dark gray-brown, oblong-clavate, 2.5–3.5 × 1.2–1.5 mm (l/w: (1.7–)2.1–3.5), apex round, usually glabrous; ribs (4–)5, broadly rounded, smooth, edges sharp and slightly overhanging sulcus; sulci about as wide as base of ribs, smooth, not papillate (very rarely minutely and sparsely puberulent). |
2n | = ca. 52. |
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Boerhavia triquetra |
Boerhavia linearifolia |
|
Phenology | Flowering summer–late fall. | Flowering mid spring–late summer. |
Habitat | Sandy or gravelly areas in deserts | Open, calcareous soils or rock in arid grasslands or shrublands |
Elevation | [0-]100-300 m ([0-]300-1000 ft) | 400-1700 m (1300-5600 ft) |
Distribution |
AZ; CA; Mexico (Baja California, Baja California Sur, Sonora)
|
NM; TX; Mexico (Chihuahua, Coahuila) |
Discussion | Boerhavia triquetra, very narrowly defined here by a predominance of 3- and 4-ribbed fruits, is uncommon in the flora, known to occur only on low deserts near the Colorado River and southward around the Sea of Cortez. The species has few- or 1-flowered terminal inflorescences. Its relationship to, and distinctness from, B. intermedia, needs clarification. In the area of the type locality, Bahia de los Angeles, on the east coast of Baja California, some plants have only 3- and 4-winged fruits, whereas most will have 3-, 4-, and some 5-winged fruits. The five-winged fruits do not differ from those of B. intermedia, a species common on the peninsula. Intergradient plants occur on the peninsula and on islands in the gulf. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Boerhavia linearifolia has been reported from Alabama, but no documenting specimen has been seen. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 4, p. 24. | FNA vol. 4, p. 22. |
Parent taxa | Nyctaginaceae > Boerhavia | Nyctaginaceae > Boerhavia |
Sibling taxa | ||
Synonyms | B. lindheimeri, B. tenuifolia | |
Name authority | S. Watson: Proc. Amer. Acad. Arts 24: 69. (1889) | A. Gray: Amer. J. Sci. Arts, ser. 2, 15: 322. (1853) |
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