Boerhavia triquetra |
Boerhavia ciliata |
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creeping sticky stem, slender spiderling |
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Habit | Herbs, annual; taproot tapered, soft or ± woody. | Herbs, perennial, slightly woody at base; taproot long, ropelike, ± woody. |
Stems | erect or ascending, occasionally decumbent, moderately or profusely branched, primarily distally, 1–5 dm, minutely puberulent with bent hairs basally, glabrous or minutely puberulent distally. |
decumbent to ascending, sparingly branched throughout, 2–7 dm, with minute, crinkled hairs and spreading long hairs, sometimes also glandular basally, glabrous distally. |
Leaves | mostly in basal 1/2 of plant; larger leaves with petiole 4–8[–15] mm, blade broadly ovate or oval to lanceolate, 10–21[–35] × 6–8[–13] mm (distal leaves smaller, proportionately narrower), adaxial surface usually glabrous, rarely minutely puberulent, abaxial surface paler than adaxial, glabrous, neither surface punctate, base round [obtuse], margins shallowly sinuate, apex acute, obtuse, or round. |
mostly in basal 1/2 of plant; larger leaves with petiole 2–24 mm, blade ovate to orbiculate, occasionally wider than long, 10–25 × 6–22 mm (distal leaves smaller, proportionately narrower), adaxial surface glabrate to sparsely pilose, especially on veins, abaxial surface much paler than adaxial, glabrous or moderately pilose, especially on veins, neither surface punctate, base round to cordate, margins entire or sinuate, apex obtuse to round. |
Inflorescences | terminal or axillary, forked ca. 3–6 times ± evenly (or clearly unevenly), diffuse, with sticky internodal bands; branches usually terminating in umbels or flowers borne singly, usually well beyond leaves. |
axillary or terminal, forked ca. 2–4 times unequally, open, without sticky internodal bands; branches ascending-divergent, terminating in loose, 1–3-flowered, cymose clusters. |
Flowers | pedicel 0.3–4.5 mm; bracts at base of perianth quickly deciduous, 2, linear-lanceolate to lanceolate [ovate], 0.5–0.6 mm; perianth whitish to pale pink or purplish, campanulate beyond constriction, 1 mm; stamens 2–3, included or barely exserted. |
pedicel 1–6 mm; bracts at base of perianth persistent, usually 3, linear to lanceolate, 1–1.5 mm; perianth pale pink to pink, occasionally purple-red, widely funnelform distal to constriction, 2.5–3 mm; stamens 3–5, well exserted. |
Fruits | 1–7 per umbel, straw colored or gray-brown, obpyramidal, 2.5–3.1 × 1.3–1.9 mm (l/w: 1.6–2.1), base stipelike, apex round-truncate, glabrous; ribs 3–4(–5), acute or winglike, tapering to pedicel, slightly rugose or undulate near sulci; sulci 0.5–1 times as wide as base of ribs, coarsely transversely rugose, smooth or very faintly papillate. |
borne singly (2–3 in open clusters), gray-brown, oblong-clavate, 2.8–3.2 × 1.3 mm (l/w: 2.3–2.7), apex broadly conic to round, glabrous; ribs 5, round or very bluntly angled, smooth or rugose near sulci, edges sharp and slightly overhanging sulcus; sulci to 1/2 times as wide as base of ribs, not rugose, minutely papillate. |
2n | = ca. 52. |
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Boerhavia triquetra |
Boerhavia ciliata |
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Phenology | Flowering summer–late fall. | Flowering late spring-mid fall. |
Habitat | Sandy or gravelly areas in deserts | Open limestone and caliche in brush, arid grasslands |
Elevation | [0-]100-300 m ([0-]300-1000 ft) | 0-100[-2000] m (0-300[-6600] ft) |
Distribution |
AZ; CA; Mexico (Baja California, Baja California Sur, Sonora)
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TX; Mexico (San Luis Potosí, Tamaulipas) |
Discussion | Boerhavia triquetra, very narrowly defined here by a predominance of 3- and 4-ribbed fruits, is uncommon in the flora, known to occur only on low deserts near the Colorado River and southward around the Sea of Cortez. The species has few- or 1-flowered terminal inflorescences. Its relationship to, and distinctness from, B. intermedia, needs clarification. In the area of the type locality, Bahia de los Angeles, on the east coast of Baja California, some plants have only 3- and 4-winged fruits, whereas most will have 3-, 4-, and some 5-winged fruits. The five-winged fruits do not differ from those of B. intermedia, a species common on the peninsula. Intergradient plants occur on the peninsula and on islands in the gulf. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 4, p. 24. | FNA vol. 4, p. 22. |
Parent taxa | Nyctaginaceae > Boerhavia | Nyctaginaceae > Boerhavia |
Sibling taxa | ||
Synonyms | B. mathisiana | |
Name authority | S. Watson: Proc. Amer. Acad. Arts 24: 69. (1889) | Brandegee: Univ. Calif. Publ. Bot. 4: 270. (1912) |
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