Boerhavia spicata |
Boerhavia coulteri |
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creeping spiderling |
Coulter's spiderling |
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Habit | Herbs, annual; taproot tapered, soft or ± woody. | Herbs, annual; taproot tapered, soft or ± woody. | ||||
Stems | erect or ascending, sparingly branched throughout, 30–70 dm, densely glandular-villous, or glandular-puberulent, with spreading, nonglandular hairs basally, glabrous distally. |
erect to decumbent-ascending, sparingly to profusely branched throughout, 2–8(–15) dm, usually minutely puberulent, often also with long, spreading hairs, occasionally also glandular basally, glabrous distally. |
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Leaves | mostly in basal 1/2; larger leaves with petiole 10–30 mm, blade oval, oblong, ovate, or ± triangular, 18–45 × 13–30 mm (distal leaves usually smaller, sometimes longer, proportionately narrower), adaxial surface lightly to densely glandular-pubescent, abaxial surface paler than adaxial, lightly to densely glandular-pubescent, neither surface punctate or both minutely punctate with clusters of brown cells, base truncate, round, or obtuse, margins sinuate, sometimes crisped, apex round to obtuse, rarely acute. |
mostly in basal 1/2; larger leaves with petiole 5–30 mm, blade lanceolate, ovate, oval, deltate-ovate, or narrowly deltate, 10–50 × 6–32 mm (distal leaves usually shorter, proportionately narrower), adaxial surface usually glabrous, sometimes sparsely hirtellous, abaxial surface paler than adaxial, usually glabrous, sometimes sparsely puberulent or with few coarse hairs, usually neither surface punctate, base acute, obtuse, or round, margins sinuate, often crisped, apex acute, obtuse, or rounded. |
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Inflorescences | terminal and axillary, branched 1–4 times unequally, with sticky internodal bands; branches strongly ascending, terminating in spicate or racemose flower clusters, axis 10–55 mm. |
terminal and axillary, forked ca. 2–5 times unevenly, usually with sticky internodal bands; branches strongly ascending, terminating in spicate or racemose flower clusters, axis 15–60 mm. |
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Flowers | pedicel 0.4–2.3[–3.7] mm; bracts at base of perianth usually soon deciduous, usually 2, narrowly to broadly lanceolate, 0.7–1.8 mm, apex often acuminate; perianth white to pale pink, campanulate distal to constriction, 1–1.3 mm; stamens (2–)3, slightly exserted or included. |
pedicel 0.1–1.6 mm; bracts at base of perianth soon deciduous, 1–2, usually lanceolate, lance-acuminate, or linear-lanceolate, rarely ovate, 0.4–1 mm; perianth white to pale pink, campanulate distal to constriction, 0.7–2 mm; stamens (1–)2–3(–4), included or slightly exserted. |
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Fruits | 5–33 per cluster, usually overlapping or 2–4 in group separated by small gap from next group, straw colored to grayish or reddish tan, broadly obovoid, 1.9–2.4(–2.8) × 1.1–1.3 mm (l/w: 1.7–2.1[–2.3]), apex rounded, glabrous; ribs 5, obtuse-rounded to obtuse, often with low winglike ridge, slightly rugose near sulci; sulci (0.2–)0.5 times as wide as base of ribs, slightly rugose, not papillate. |
4–20(–22) per cluster, remotely spaced or overlapped by 1–100% of their length, or 2–4 in group, separated by small gap from next group and with distal spikelets overlapping, straw colored to pale red-brown, narrowly obovoid to narrowly obpyramidal, 2–3.2(–3.6) × 0.9–1.1(–1.4) mm (l/w: [1.9–]2.1–3.1[–3.3]), apex rounded to bluntly conic-truncate, or truncate, glabrous; ribs 5, obtuse or round-obtuse, often with sharp ridges, slightly rugose near sulci; sulci 0.1–0.3 times as wide as base of ribs, smooth or slightly rugose, not papillate. |
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2n | = ca. 52. |
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Boerhavia spicata |
Boerhavia coulteri |
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Phenology | Flowering late summer–early fall. | |||||
Habitat | Sandy or rocky soils in open, arid grasslands, among open shrubs or mesquite and acacia woodlands [tropical deciduous forests] | |||||
Elevation | [100-]700-1800 m ([300-]2300-5900 ft) | |||||
Distribution |
AZ; NM; TX; Mexico
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AZ; CA; NM; TX; UT; n Mexico
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Discussion | C. F. Reed (1969) and subsequent authors have included one or more of Boerhavia coulteri, B. torreyana, and B. watsoni as synonymous with B. spicata. Even when those taxa are removed, B. spicata remains a variable species, widespread at low to middle elevations in southwestern North America, and distinguished by its mostly overlapping, obovoid fruits with rather open sulci, and the glandular pubescence on basal parts of the plant. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 2 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 4, p. 25. | FNA vol. 4, p. 26. | ||||
Parent taxa | Nyctaginaceae > Boerhavia | Nyctaginaceae > Boerhavia | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Senkenbergia coulteri | |||||
Name authority | Choisy: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 13(2): 456. (1849) | (Hooker f.) S. Watson: Proc. Amer. Acad. Arts 24: 70. (1889) | ||||
Web links |