Boerhavia spicata |
Boerhavia ciliata |
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creeping spiderling |
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Habit | Herbs, annual; taproot tapered, soft or ± woody. | Herbs, perennial, slightly woody at base; taproot long, ropelike, ± woody. |
Stems | erect or ascending, sparingly branched throughout, 30–70 dm, densely glandular-villous, or glandular-puberulent, with spreading, nonglandular hairs basally, glabrous distally. |
decumbent to ascending, sparingly branched throughout, 2–7 dm, with minute, crinkled hairs and spreading long hairs, sometimes also glandular basally, glabrous distally. |
Leaves | mostly in basal 1/2; larger leaves with petiole 10–30 mm, blade oval, oblong, ovate, or ± triangular, 18–45 × 13–30 mm (distal leaves usually smaller, sometimes longer, proportionately narrower), adaxial surface lightly to densely glandular-pubescent, abaxial surface paler than adaxial, lightly to densely glandular-pubescent, neither surface punctate or both minutely punctate with clusters of brown cells, base truncate, round, or obtuse, margins sinuate, sometimes crisped, apex round to obtuse, rarely acute. |
mostly in basal 1/2 of plant; larger leaves with petiole 2–24 mm, blade ovate to orbiculate, occasionally wider than long, 10–25 × 6–22 mm (distal leaves smaller, proportionately narrower), adaxial surface glabrate to sparsely pilose, especially on veins, abaxial surface much paler than adaxial, glabrous or moderately pilose, especially on veins, neither surface punctate, base round to cordate, margins entire or sinuate, apex obtuse to round. |
Inflorescences | terminal and axillary, branched 1–4 times unequally, with sticky internodal bands; branches strongly ascending, terminating in spicate or racemose flower clusters, axis 10–55 mm. |
axillary or terminal, forked ca. 2–4 times unequally, open, without sticky internodal bands; branches ascending-divergent, terminating in loose, 1–3-flowered, cymose clusters. |
Flowers | pedicel 0.4–2.3[–3.7] mm; bracts at base of perianth usually soon deciduous, usually 2, narrowly to broadly lanceolate, 0.7–1.8 mm, apex often acuminate; perianth white to pale pink, campanulate distal to constriction, 1–1.3 mm; stamens (2–)3, slightly exserted or included. |
pedicel 1–6 mm; bracts at base of perianth persistent, usually 3, linear to lanceolate, 1–1.5 mm; perianth pale pink to pink, occasionally purple-red, widely funnelform distal to constriction, 2.5–3 mm; stamens 3–5, well exserted. |
Fruits | 5–33 per cluster, usually overlapping or 2–4 in group separated by small gap from next group, straw colored to grayish or reddish tan, broadly obovoid, 1.9–2.4(–2.8) × 1.1–1.3 mm (l/w: 1.7–2.1[–2.3]), apex rounded, glabrous; ribs 5, obtuse-rounded to obtuse, often with low winglike ridge, slightly rugose near sulci; sulci (0.2–)0.5 times as wide as base of ribs, slightly rugose, not papillate. |
borne singly (2–3 in open clusters), gray-brown, oblong-clavate, 2.8–3.2 × 1.3 mm (l/w: 2.3–2.7), apex broadly conic to round, glabrous; ribs 5, round or very bluntly angled, smooth or rugose near sulci, edges sharp and slightly overhanging sulcus; sulci to 1/2 times as wide as base of ribs, not rugose, minutely papillate. |
2n | = ca. 52. |
= ca. 52. |
Boerhavia spicata |
Boerhavia ciliata |
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Phenology | Flowering late summer–early fall. | Flowering late spring-mid fall. |
Habitat | Sandy or rocky soils in open, arid grasslands, among open shrubs or mesquite and acacia woodlands [tropical deciduous forests] | Open limestone and caliche in brush, arid grasslands |
Elevation | [100-]700-1800 m ([300-]2300-5900 ft) | 0-100[-2000] m (0-300[-6600] ft) |
Distribution |
AZ; NM; TX; Mexico
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TX; Mexico (San Luis Potosí, Tamaulipas) |
Discussion | C. F. Reed (1969) and subsequent authors have included one or more of Boerhavia coulteri, B. torreyana, and B. watsoni as synonymous with B. spicata. Even when those taxa are removed, B. spicata remains a variable species, widespread at low to middle elevations in southwestern North America, and distinguished by its mostly overlapping, obovoid fruits with rather open sulci, and the glandular pubescence on basal parts of the plant. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 4, p. 25. | FNA vol. 4, p. 22. |
Parent taxa | Nyctaginaceae > Boerhavia | Nyctaginaceae > Boerhavia |
Sibling taxa | ||
Synonyms | B. mathisiana | |
Name authority | Choisy: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 13(2): 456. (1849) | Brandegee: Univ. Calif. Publ. Bot. 4: 270. (1912) |
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