Boerhavia erecta |
Boerhavia pterocarpa |
|
---|---|---|
erect boerhavia, erect spiderling |
Apache Pass spiderling, wing-fruit spiderling |
|
Habit | Herbs, annual [slightly woody at base]; taproot tapered, soft or ± woody. | Herbs, annual; taproot tapered, soft or ± woody. |
Stems | usually erect, sometimes decumbent, profusely branched primarily distally, 2–12 dm, minutely puberulent with bent hairs basally, usually glabrous, rarely sparsely puberulent distally. |
procumbent or decumbent to ascending, sparingly branched throughout, 1–4 dm, minutely puberulent with bent hairs throughout. |
Leaves | mostly in basal 1/2 of plant; larger leaves with petiole 6–40(–55) mm, blade broadly rhombic-ovate, triangular-ovate, ovate, oval, or lanceolate, 20–50(–80) × 10–45 mm (distal leaves smaller, proportionately narrower), adaxial surface usually glabrous, sometimes minutely puberulent, usually minutely punctate, abaxial surface slightly paler than adaxial, usually glabrous, sometimes minutely puberulent, usually punctate with small patches of small brown cells, base obtuse to round, margins entire or sinuate, apex usually acute, less often obtuse or rounded. |
± throughout; larger leaves with petiole 3–12 mm, blade rhombic-ovate to ovate or lanceolate, 15–25 × 9–15 mm (distal leaves smaller, proportionately narrower), adaxial surface glabrous or sparsely puberulent, abaxial surface paler than adaxial, glabrous, neither surface punctate, base round to obtuse, margins entire or slightly sinuate, apex acute. |
Inflorescences | terminal, forked ca. 4–6 times ± evenly, diffuse, usually with sticky internodal bands; branches strongly ascending, terminating in irregular umbellate or subracemose clusters of flowers, not all pedicels attaching at same point (flowers occasionally borne singly). |
terminal or axillary, without sticky internodal bands; peduncle, 1–3 cm, bearing small capitate clusters of flowers. |
Flowers | pedicel (0–)0.3–2.5(–5) mm; bracts at base of perianth deciduous, usually 2, narrowly to broadly lanceolate, 0.5–1 mm, apex often acuminate; perianth whitish, usually tinged with pink or purple [bright pink] between lobes and in tube, campanulate beyond constriction, 1–1.5 mm; stamens 2–4, slightly exserted. |
pedicel 0.3–0.6 mm; bracts at base of perianth quickly deciduous, lance-acuminate, 0.4–0.7 mm; perianth white to pale pinkish, campanulate distal to constriction, 1–1.5 mm; stamens 2, included or barely exserted. |
Fruits | 1–11 per cluster, pale greenish to straw colored or tan, narrowly obconic, (2.7–)3–3.5[–4] × 1.2–1.5 mm (l/w: (2–)2.3–3.2), apex truncate or broadly low conic, glabrous; ribs 5, acute, slightly rugose adjacent to sulci; sulci 0.5–1 times as wide as base of ribs, slightly to prominently coarsely transverse rugose, not papillate. |
2–8 per cluster, pale green to straw colored, broadly obpyramidal, base tapered to stipelike above pedicel, 2.9–3.4 × 2.8–3.2 mm (l/w: 1–1.4), apex truncate, glabrous; ribs 3–4, winglike, smooth; sulci 3–4 times as wide as base of ribs, coarsely transversely rugose, not papillate. |
Boerhavia erecta |
Boerhavia pterocarpa |
|
Phenology | Flowering early summer-mid fall. | Flowering late summer. |
Habitat | Disturbed areas, gardens, road and railroad rights-of-way, stream beds | Sandy loam to clay soils, disturbed areas, occasionally a weed in ornamental beds |
Elevation | 0-1700 m [probably much higher in tropics] (0-5600 ft [probably much higher in tropics]) | 700-1200 m (2300-3900 ft) |
Distribution |
AL; AR; AZ; FL; GA; LA; MD; MO; MS; NC; NM; OK; SC; TN; TX; Mexico; Central America; South America; West Indies [Widely introduced throughout the tropics and warm-temperate regions]
|
AZ; Mexico (Sonora)
|
Discussion | Boerhavia erecta occasionally forms mixed populations with B. intermedia without apparent intergradation. Rarely, some specimens seem to combine features of either species, particularly with regard to inflorescence structure. This is especially so in Sonora, Mexico, and in parts of the Sonoran Desert in Arizona. The two species bloom simultaneously and are visited by small insects. Given the presumed close relationship and weedy habitats of each, hybridization seems possible. Usually, the two species can be distinguished by the differences in fruit length, the appearance of a crownlike apex of the nearly mature fruits of B. erecta (apex of ridges slightly expanded, apex of fruit slightly conic), and the more precisely constructed terminal umbels of B. intermedia. Both species, particularly B. intermedia, may produce entire inflorescences with branches terminating in single flowers. R. E. Woodson Jr. and H. J. Kidd (1961) suggested that B. erecta hybridizes with the perennial B. diffusa. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
|
Source | FNA vol. 4, p. 22. | FNA vol. 4, p. 24. |
Parent taxa | Nyctaginaceae > Boerhavia | Nyctaginaceae > Boerhavia |
Sibling taxa | ||
Name authority | Linnaeus: Sp. Pl. 1: 3. (1753) | S. Watson: Proc. Amer. Acad. Arts 17: 376. (1882) |
Web links |