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erect boerhavia, erect spiderling

Tucson Mountain spiderling

Habit Herbs, annual [slightly woody at base]; taproot tapered, soft or ± woody. Herbs, annual; taproot tapered, soft or ± woody.
Stems

usually erect, sometimes decumbent, profusely branched primarily distally, 2–12 dm, minutely puberulent with bent hairs basally, usually glabrous, rarely sparsely puberulent distally.

usually erect or ascending, occasionally decumbent, branched several times, primarily distally, 3–6 dm, minutely puberulent with bent hairs and sometimes also glandular hairs in basal portions; sparsely to moderately puberulent distally.

Leaves

mostly in basal 1/2 of plant;

larger leaves with petiole 6–40(–55) mm, blade broadly rhombic-ovate, triangular-ovate, ovate, oval, or lanceolate, 20–50(–80) × 10–45 mm (distal leaves smaller, proportionately narrower), adaxial surface usually glabrous, sometimes minutely puberulent, usually minutely punctate, abaxial surface slightly paler than adaxial, usually glabrous, sometimes minutely puberulent, usually punctate with small patches of small brown cells, base obtuse to round, margins entire or sinuate, apex usually acute, less often obtuse or rounded.

mostly in basal 1/2 of plant;

larger leaves with petiole 12–17 mm, blade oblong-lanceolate, 25–50 × 10–17 mm (distal leaves smaller, sometimes longer, proportionately narrower), adaxial surface glabrous or very sparsely puberulent, often punctate, abaxial surface paler than adaxial, glabrous or very sparsely puberulent, usually punctate with small dark brown cells, base round, obtuse, or truncate, margins entire or slightly and irregularly crenulate, apex broadly to narrowly acute.

Inflorescences

terminal, forked ca. 4–6 times ± evenly, diffuse, usually with sticky internodal bands;

branches strongly ascending, terminating in irregular umbellate or subracemose clusters of flowers, not all pedicels attaching at same point (flowers occasionally borne singly).

terminal, forked ± unevenly ca. 3–4 times, with sticky internodal bands;

branches strongly ascending, terminating in umbels (rarely flowers borne singly).

Flowers

pedicel (0–)0.3–2.5(–5) mm;

bracts at base of perianth deciduous, usually 2, narrowly to broadly lanceolate, 0.5–1 mm, apex often acuminate;

perianth whitish, usually tinged with pink or purple [bright pink] between lobes and in tube, campanulate beyond constriction, 1–1.5 mm;

stamens 2–4, slightly exserted.

pedicel 1–2 mm;

bracts at base of perianth quickly deciduous, 2, lanceolate, 0.6–1.5 mm;

perianth whitish to pale pink, campanulate distal to constriction, 1–1.5 mm;

stamens 3–4, barely exserted.

Fruits

1–11 per cluster, pale greenish to straw colored or tan, narrowly obconic, (2.7–)3–3.5[–4] × 1.2–1.5 mm (l/w: (2–)2.3–3.2), apex truncate or broadly low conic, glabrous;

ribs 5, acute, slightly rugose adjacent to sulci;

sulci 0.5–1 times as wide as base of ribs, slightly to prominently coarsely transverse rugose, not papillate.

(1–)3–14 per umbel, straw colored at maturity, broadly obconic, emarginate in profile (wings extended beyond apex of body), 3.2–3.8 × 2.2–2.6 mm (l/w: 1.3–1.5), glabrous;

ribs 5, winglike, smooth;

sulci 1–2 times as wide as base of ribs, smooth or very slightly rugose, not papillate.

Boerhavia erecta

Boerhavia megaptera

Phenology Flowering early summer-mid fall. Flowering summer–fall.
Habitat Disturbed areas, gardens, road and railroad rights-of-way, stream beds Rocky ground, among desert shrubs or trees
Elevation 0-1700 m [probably much higher in tropics] (0-5600 ft [probably much higher in tropics]) 700-1300 m (2300-4300 ft)
Distribution
from FNA
AL; AR; AZ; FL; GA; LA; MD; MO; MS; NC; NM; OK; SC; TN; TX; Mexico; Central America; South America; West Indies [Widely introduced throughout the tropics and warm-temperate regions]
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ
[BONAP county map]
Discussion

Boerhavia erecta occasionally forms mixed populations with B. intermedia without apparent intergradation. Rarely, some specimens seem to combine features of either species, particularly with regard to inflorescence structure. This is especially so in Sonora, Mexico, and in parts of the Sonoran Desert in Arizona. The two species bloom simultaneously and are visited by small insects. Given the presumed close relationship and weedy habitats of each, hybridization seems possible. Usually, the two species can be distinguished by the differences in fruit length, the appearance of a crownlike apex of the nearly mature fruits of B. erecta (apex of ridges slightly expanded, apex of fruit slightly conic), and the more precisely constructed terminal umbels of B. intermedia. Both species, particularly B. intermedia, may produce entire inflorescences with branches terminating in single flowers. R. E. Woodson Jr. and H. J. Kidd (1961) suggested that B. erecta hybridizes with the perennial B. diffusa.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Boerhavia megaptera is often sympatric with B. intermedia.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 4, p. 22. FNA vol. 4, p. 24.
Parent taxa Nyctaginaceae > Boerhavia Nyctaginaceae > Boerhavia
Sibling taxa
B. anisophylla, B. ciliata, B. coccinea, B. coulteri, B. diffusa, B. gracillima, B. intermedia, B. linearifolia, B. megaptera, B. pterocarpa, B. purpurascens, B. spicata, B. torreyana, B. triquetra, B. wrightii
B. anisophylla, B. ciliata, B. coccinea, B. coulteri, B. diffusa, B. erecta, B. gracillima, B. intermedia, B. linearifolia, B. pterocarpa, B. purpurascens, B. spicata, B. torreyana, B. triquetra, B. wrightii
Name authority Linnaeus: Sp. Pl. 1: 3. (1753) Standley: Contr. U.S. Natl. Herb. 12: 379. (1909)
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