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erect boerhavia, erect spiderling

wineflower

Habit Herbs, annual [slightly woody at base]; taproot tapered, soft or ± woody. Herbs, perennial, often somewhat woody at base; taproot long and ropelike, woody.
Stems

usually erect, sometimes decumbent, profusely branched primarily distally, 2–12 dm, minutely puberulent with bent hairs basally, usually glabrous, rarely sparsely puberulent distally.

decumbent to ascending, sparingly branched throughout, 2–8 dm, densely glandular-pubescent or pubescent with minute curled hairs, sometimes also with sparse long-spreading hairs basally, usually glabrous sometimes glandular-puberulent distally.

Leaves

mostly in basal 1/2 of plant;

larger leaves with petiole 6–40(–55) mm, blade broadly rhombic-ovate, triangular-ovate, ovate, oval, or lanceolate, 20–50(–80) × 10–45 mm (distal leaves smaller, proportionately narrower), adaxial surface usually glabrous, sometimes minutely puberulent, usually minutely punctate, abaxial surface slightly paler than adaxial, usually glabrous, sometimes minutely puberulent, usually punctate with small patches of small brown cells, base obtuse to round, margins entire or sinuate, apex usually acute, less often obtuse or rounded.

mostly in basal 1/2 of plant;

larger leaves with petiole 4–22 mm, blade triangular-ovate to ovate or suborbiculate, 20–40 × 15–32 mm (distal leaves smaller, proportionately narrower), adaxial surface glabrate to densely spreading-puberulent, rarely lightly punctate, abaxial surface paler than adaxial, glabrate to moderately glandular-puberulent, usually punctate with small patches of large gray cells, base round, truncate, or cordate, usually oblique, margins entire or sinuate, apex obtuse to round.

Inflorescences

terminal, forked ca. 4–6 times ± evenly, diffuse, usually with sticky internodal bands;

branches strongly ascending, terminating in irregular umbellate or subracemose clusters of flowers, not all pedicels attaching at same point (flowers occasionally borne singly).

axillary or terminal, forked ca. 3–5 times unequally, open, without sticky internodal bands;

branches strongly ascending, terminating in loose, cymose, 1–3-flowered clusters.

Flowers

pedicel (0–)0.3–2.5(–5) mm;

bracts at base of perianth deciduous, usually 2, narrowly to broadly lanceolate, 0.5–1 mm, apex often acuminate;

perianth whitish, usually tinged with pink or purple [bright pink] between lobes and in tube, campanulate beyond constriction, 1–1.5 mm;

stamens 2–4, slightly exserted.

pedicel 0.5–5 mm;

bracts at base of perianth persistent, 2–3, narrowly to broadly lanceolate, 1–2 mm, apex often acuminate;

perianth purplish pink to pink (white), widely funnelform beyond constriction, 5–10 mm;

stamens (4–)5–8, well exserted.

Fruits

1–11 per cluster, pale greenish to straw colored or tan, narrowly obconic, (2.7–)3–3.5[–4] × 1.2–1.5 mm (l/w: (2–)2.3–3.2), apex truncate or broadly low conic, glabrous;

ribs 5, acute, slightly rugose adjacent to sulci;

sulci 0.5–1 times as wide as base of ribs, slightly to prominently coarsely transverse rugose, not papillate.

borne singly (or 2–3 in open clusters), gray-brown, oblong-obovoid, 2.7–3.6 × 1.3–1.6 mm (l/w: [1.7–]2.1–2.6), apex rounded to broad-conic, glabrous;

ribs 5, round or bluntly angled, not overhanging sulci, smooth;

sulci 1–2 times as wide as base of ribs, not rugose (rarely deeply transversely rugose), minutely papillate.

Boerhavia erecta

Boerhavia anisophylla

Phenology Flowering early summer-mid fall. Flowering mid summer–early fall.
Habitat Disturbed areas, gardens, road and railroad rights-of-way, stream beds Limestone banks and calcareous gravel in arid shrublands
Elevation 0-1700 m [probably much higher in tropics] (0-5600 ft [probably much higher in tropics]) 1000-1300[-2300] m (3300-4300[-7500] ft)
Distribution
from FNA
AL; AR; AZ; FL; GA; LA; MD; MO; MS; NC; NM; OK; SC; TN; TX; Mexico; Central America; South America; West Indies [Widely introduced throughout the tropics and warm-temperate regions]
[WildflowerSearch map]
[BONAP county map]
from FNA
TX; Mexico (Chihuahua, Coahuila, Durango, Nuevo León, Zacatecas)
[BONAP county map]
Discussion

Boerhavia erecta occasionally forms mixed populations with B. intermedia without apparent intergradation. Rarely, some specimens seem to combine features of either species, particularly with regard to inflorescence structure. This is especially so in Sonora, Mexico, and in parts of the Sonoran Desert in Arizona. The two species bloom simultaneously and are visited by small insects. Given the presumed close relationship and weedy habitats of each, hybridization seems possible. Usually, the two species can be distinguished by the differences in fruit length, the appearance of a crownlike apex of the nearly mature fruits of B. erecta (apex of ridges slightly expanded, apex of fruit slightly conic), and the more precisely constructed terminal umbels of B. intermedia. Both species, particularly B. intermedia, may produce entire inflorescences with branches terminating in single flowers. R. E. Woodson Jr. and H. J. Kidd (1961) suggested that B. erecta hybridizes with the perennial B. diffusa.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 4, p. 22. FNA vol. 4, p. 21.
Parent taxa Nyctaginaceae > Boerhavia Nyctaginaceae > Boerhavia
Sibling taxa
B. anisophylla, B. ciliata, B. coccinea, B. coulteri, B. diffusa, B. gracillima, B. intermedia, B. linearifolia, B. megaptera, B. pterocarpa, B. purpurascens, B. spicata, B. torreyana, B. triquetra, B. wrightii
B. ciliata, B. coccinea, B. coulteri, B. diffusa, B. erecta, B. gracillima, B. intermedia, B. linearifolia, B. megaptera, B. pterocarpa, B. purpurascens, B. spicata, B. torreyana, B. triquetra, B. wrightii
Name authority Linnaeus: Sp. Pl. 1: 3. (1753) Torrey: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 171. (1859)
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