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red spiderling, spreading hogweed

narrowleaf spiderling

Habit Herbs, perennial [rarely appearing annual], sometimes slightly woody at base; taproot fusiform, woody. Herbs, perennial, sometimes slightly woody at base; taproot long, ropelike, ± woody.
Stems

decumbent, ascending, or erect, usually profusely branched throughout, 3–10 dm, glabrous or minutely pubescent basally, glabrous or sparsely pubescent distally.

usually erect or ascending, occasionally decumbent, sparingly branched throughout, 2–5(–9) dm, hirsute, puberulent, or glandular basally, hair types often mixed, glandular-puberulent, glabrous, or occasionally puberulent and sparsely hispid distally.

Leaves

mostly in basal 1/2 of plant;

larger leaves with petiole 10–30 mm, blade broadly lanceolate, ovate, or broadly ovate, occasionally ± round, 20–60 × 15–50 mm (distal leaves smaller, often proportionally narrower), base truncate, broadly cuneate, round, or shallowly cordate, often oblique, margins sinuate, apex obtuse to round, abaxial surface paler than adaxial, glabrous or minutely puberulent, sometimes glandular, usually with few large multicellular hairs on veins, adaxial surface glabrous, rarely sparsely puberulent, neither surface punctate.

mostly in basal 2/3 of plant;

larger leaves with petiole 1–5 mm, blade lanceolate to linear, rarely ovate, 15–35 × 1–15 mm (distal leaves usually longer, narrower), adaxial surface glabrous or sparsely hispid, occasionally glandular-pubescent, sometimes punctate, abaxial surface usually paler than adaxial, glabrous or sparsely hispid, occasionally glandular-pubescent, usually punctate with small patches of large gray to dark brown cells, base obtuse, truncate, or rounded, margins entire and revolute, apex usually acute, sometimes obtuse or rounded.

Inflorescences

terminal, forked ca. 3–6 times ± equally, diffuse, occasionally with sticky internodal bands;

branches divergent, terminating in compact subumbellate or capitate, 2–5-flowered clusters.

terminal and axillary, forked ca. 2–4 times unequally, open, without sticky internodal bands;

branches strongly ascending, terminating in loose, 1–few-flowered, cymose clusters.

Flowers

pedicel shorter than 0.5 mm;

bract at base of perianth quickly deciduous, 1, lanceolate, 0.8–1 mm;

perianth purplish red to reddish pink or nearly white, campanulate beyond constriction, 1–1.5 mm;

stamens 2–3, included or barely exserted.

pedicel 1–2(–5) mm;

bracts at base of perianth persistent, 3, lanceolate to ovate-acuminate, 1–2 mm;

perianth purplish pink, widely funnelform beyond constriction, 4–7 mm;

stamens 5(–6), well exserted.

Fruits

(1–)2–5(–9) per cluster, gray-brown to brown, obpyramidal, (3–)3.5–4.5 × 1–1.2 mm (l/w: 2.8–4.1), apex broadly conic, with sparse or moderately dense stipitate-glandular hairs on ribs, less densely pubescent or glabrous in sulci [rarely entirely glabrous];

ribs 5, round, smooth;

sulci ± 2–2.5 times as wide as base of ribs, not rugose, not papillate.

borne singly (or 2–3 in open clusters), dark gray-brown, oblong-clavate, 2.5–3.5 × 1.2–1.5 mm (l/w: (1.7–)2.1–3.5), apex round, usually glabrous;

ribs (4–)5, broadly rounded, smooth, edges sharp and slightly overhanging sulcus;

sulci about as wide as base of ribs, smooth, not papillate (very rarely minutely and sparsely puberulent).

2n

= 26, 52, 54, 116.

= ca. 52.

Boerhavia diffusa

Boerhavia linearifolia

Phenology Flowering mid winter–early fall [year-round]. Flowering mid spring–late summer.
Habitat Disturbed areas, waste places, roadsides, dry pinelands, among scrub on tropical reefs Open, calcareous soils or rock in arid grasslands or shrublands
Elevation 0-50[-1800] m (0-200[-5900] ft) 400-1700 m (1300-5600 ft)
Distribution
from FNA
FL; GA; SC; Mexico; Central America; South America; West Indies; Asia; Africa; Indian Ocean Islands; Pacific Islands; Australia
[WildflowerSearch map]
[BONAP county map]
from FNA
NM; TX; Mexico (Chihuahua, Coahuila)
[BONAP county map]
Discussion

Boerhavia diffusa belongs to a pantropical complex whose distribution, because of differing taxonomic treatments, is not precisely discernable. In North America, the West Indies, and elsewhere, the complex has been treated as having one (e.g., R. W. Long and O. Lakela 1971; R. P. Wunderlin 1998) or two species (E. A. Kellogg 1988; J. K. Small 1913c; P. C. Standley 1918; C. Whitehouse 1996). In the latter situation various binomials have been used. The abrupt bend in the ribs near the fruit apex, making a broadly conic apex, seems to be a useful character for distinguishing most B. diffusa. The shape of the apex, the sparse pubescence of the fruit, the few fruits in individual terminal clusters, and the open, ± leafless inflorescence may have led R. E. Woodson Jr. and H. J. Kidd (1961) to suspect hybridization of this complex with the annual B. erecta. Hybridization is plausible (R. Spellenberg 2000), but clearly intermediate plants are not known.

The leaves are sometimes used as a vegetable (C. Whitehouse 1996). Extracts from roots are used to prepare an expectorant, a diuretic, and a laxative, and in treating asthma (S. P. Ambasta 1986).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Boerhavia linearifolia has been reported from Alabama, but no documenting specimen has been seen.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 4, p. 19. FNA vol. 4, p. 22.
Parent taxa Nyctaginaceae > Boerhavia Nyctaginaceae > Boerhavia
Sibling taxa
B. anisophylla, B. ciliata, B. coccinea, B. coulteri, B. erecta, B. gracillima, B. intermedia, B. linearifolia, B. megaptera, B. pterocarpa, B. purpurascens, B. spicata, B. torreyana, B. triquetra, B. wrightii
B. anisophylla, B. ciliata, B. coccinea, B. coulteri, B. diffusa, B. erecta, B. gracillima, B. intermedia, B. megaptera, B. pterocarpa, B. purpurascens, B. spicata, B. torreyana, B. triquetra, B. wrightii
Synonyms B. lindheimeri, B. tenuifolia
Name authority Linnaeus: Sp. Pl. 1: 3. (1753) A. Gray: Amer. J. Sci. Arts, ser. 2, 15: 322. (1853)
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