Boerhavia diffusa
Boerhavia anisophylla
red spiderling, spreading hogweed
wineflower
decumbent, ascending, or erect, usually profusely branched throughout, 3–10 dm, glabrous or minutely pubescent basally, glabrous or sparsely pubescent distally.
decumbent to ascending, sparingly branched throughout, 2–8 dm, densely glandular-pubescent or pubescent with minute curled hairs, sometimes also with sparse long-spreading hairs basally, usually glabrous sometimes glandular-puberulent distally.
mostly in basal 1/2 of plant;
larger leaves with petiole 10–30 mm, blade broadly lanceolate, ovate, or broadly ovate, occasionally ± round, 20–60 × 15–50 mm (distal leaves smaller, often proportionally narrower), base truncate, broadly cuneate, round, or shallowly cordate, often oblique, margins sinuate, apex obtuse to round, abaxial surface paler than adaxial, glabrous or minutely puberulent, sometimes glandular, usually with few large multicellular hairs on veins, adaxial surface glabrous, rarely sparsely puberulent, neither surface punctate.
mostly in basal 1/2 of plant;
larger leaves with petiole 4–22 mm, blade triangular-ovate to ovate or suborbiculate, 20–40 × 15–32 mm (distal leaves smaller, proportionately narrower), adaxial surface glabrate to densely spreading-puberulent, rarely lightly punctate, abaxial surface paler than adaxial, glabrate to moderately glandular-puberulent, usually punctate with small patches of large gray cells, base round, truncate, or cordate, usually oblique, margins entire or sinuate, apex obtuse to round.
terminal, forked ca. 3–6 times ± equally, diffuse, occasionally with sticky internodal bands;
branches divergent, terminating in compact subumbellate or capitate, 2–5-flowered clusters.
axillary or terminal, forked ca. 3–5 times unequally, open, without sticky internodal bands;
branches strongly ascending, terminating in loose, cymose, 1–3-flowered clusters.
pedicel shorter than 0.5 mm;
bract at base of perianth quickly deciduous, 1, lanceolate, 0.8–1 mm;
perianth purplish red to reddish pink or nearly white, campanulate beyond constriction, 1–1.5 mm;
stamens 2–3, included or barely exserted.
pedicel 0.5–5 mm;
bracts at base of perianth persistent, 2–3, narrowly to broadly lanceolate, 1–2 mm, apex often acuminate;
perianth purplish pink to pink (white), widely funnelform beyond constriction, 5–10 mm;
stamens (4–)5–8, well exserted.
(1–)2–5(–9) per cluster, gray-brown to brown, obpyramidal, (3–)3.5–4.5 × 1–1.2 mm (l/w: 2.8–4.1), apex broadly conic, with sparse or moderately dense stipitate-glandular hairs on ribs, less densely pubescent or glabrous in sulci [rarely entirely glabrous];
ribs 5, round, smooth;
sulci ± 2–2.5 times as wide as base of ribs, not rugose, not papillate.
borne singly (or 2–3 in open clusters), gray-brown, oblong-obovoid, 2.7–3.6 × 1.3–1.6 mm (l/w: [1.7–]2.1–2.6), apex rounded to broad-conic, glabrous;
ribs 5, round or bluntly angled, not overhanging sulci, smooth;
sulci 1–2 times as wide as base of ribs, not rugose (rarely deeply transversely rugose), minutely papillate.
= 26, 52, 54, 116.
Boerhavia diffusa
Boerhavia anisophylla
Boerhavia diffusa belongs to a pantropical complex whose distribution, because of differing taxonomic treatments, is not precisely discernable. In North America, the West Indies, and elsewhere, the complex has been treated as having one (e.g., R. W. Long and O. Lakela 1971; R. P. Wunderlin 1998) or two species (E. A. Kellogg 1988; J. K. Small 1913c; P. C. Standley 1918; C. Whitehouse 1996). In the latter situation various binomials have been used. The abrupt bend in the ribs near the fruit apex, making a broadly conic apex, seems to be a useful character for distinguishing most B. diffusa. The shape of the apex, the sparse pubescence of the fruit, the few fruits in individual terminal clusters, and the open, ± leafless inflorescence may have led R. E. Woodson Jr. and H. J. Kidd (1961) to suspect hybridization of this complex with the annual B. erecta. Hybridization is plausible (R. Spellenberg 2000), but clearly intermediate plants are not known.
The leaves are sometimes used as a vegetable (C. Whitehouse 1996). Extracts from roots are used to prepare an expectorant, a diuretic, and a laxative, and in treating asthma (S. P. Ambasta 1986).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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