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hierba de la hormiga, hierba del cancer, red boerhaavia, red spiderling, scarlet spiderling

erect boerhavia, erect spiderling

Habit Herbs, perennial, often ± woody at base; taproot long and ropelike, woody. Herbs, annual [slightly woody at base]; taproot tapered, soft or ± woody.
Stems

prostrate to decumbent, usually profusely branched throughout, 3–15 dm, minutely pubescent, often glandular, sometimes spreading villous or hirsute in basal portions, minutely pubescent, sometimes glandular, glabrate, or glabrous distally.

usually erect, sometimes decumbent, profusely branched primarily distally, 2–12 dm, minutely puberulent with bent hairs basally, usually glabrous, rarely sparsely puberulent distally.

Leaves

usually distributed throughout plant and into much of inflorescence;

larger leaves with petiole 5–25 mm, blade broadly lanceolate, ovate, or broadly ovate, occasionally ± round, 20–70 × 10–60 mm (distal leaves smaller, often proportionally narrower), base truncate, broadly cuneate, or round, rarely cordate, margins sinuate, apex acute to obtuse or round, adaxial surface glabrous or sometimes sparsely puberulent, rarely densely glandular-pubescent, abaxial surface paler than adaxial surface, glabrous or sometimes sparsely puberulent, rarely densely glandular-pubescent, often with large multicellular hairs along veins, neither surface punctate.

mostly in basal 1/2 of plant;

larger leaves with petiole 6–40(–55) mm, blade broadly rhombic-ovate, triangular-ovate, ovate, oval, or lanceolate, 20–50(–80) × 10–45 mm (distal leaves smaller, proportionately narrower), adaxial surface usually glabrous, sometimes minutely puberulent, usually minutely punctate, abaxial surface slightly paler than adaxial, usually glabrous, sometimes minutely puberulent, usually punctate with small patches of small brown cells, base obtuse to round, margins entire or sinuate, apex usually acute, less often obtuse or rounded.

Inflorescences

axillary or terminal, forked unequally ca. 3–6 times, open, without sticky internodal bands;

branches divergent, terminating in compact subumbellate or capitate 5-flowered clusters.

terminal, forked ca. 4–6 times ± evenly, diffuse, usually with sticky internodal bands;

branches strongly ascending, terminating in irregular umbellate or subracemose clusters of flowers, not all pedicels attaching at same point (flowers occasionally borne singly).

Flowers

pedicel shorter than 0.5 mm;

bract at base of perianth usually quickly deciduous, 1, linear-lanceolate to ovate, 0.5–1 mm;

perianth maroon, or magenta (or rarely white or yellow) [pink], campanulate beyond constriction, 1–3.5 mm;

stamens 2–3, slightly exserted.

pedicel (0–)0.3–2.5(–5) mm;

bracts at base of perianth deciduous, usually 2, narrowly to broadly lanceolate, 0.5–1 mm, apex often acuminate;

perianth whitish, usually tinged with pink or purple [bright pink] between lobes and in tube, campanulate beyond constriction, 1–1.5 mm;

stamens 2–4, slightly exserted.

Fruits

(2–)5–20(–30) per cluster, gray-brown to brown, narrowly obovate and tapering at both ends or clavate, 2.6–4 × 0.9–1.2 mm (l/w: 2.7–3.5), apex rounded to rounded-conic, moderately densely to densely stipitate-glandular on ribs and in sulci;

ribs 5, rounded, smooth;

sulci 1–2.5 times as wide as base of ribs, not rugose, not papillate.

1–11 per cluster, pale greenish to straw colored or tan, narrowly obconic, (2.7–)3–3.5[–4] × 1.2–1.5 mm (l/w: (2–)2.3–3.2), apex truncate or broadly low conic, glabrous;

ribs 5, acute, slightly rugose adjacent to sulci;

sulci 0.5–1 times as wide as base of ribs, slightly to prominently coarsely transverse rugose, not papillate.

2n

= 52.

Boerhavia coccinea

Boerhavia erecta

Phenology Flowering spring–winter [year-round]. Flowering early summer-mid fall.
Habitat Roadsides, weedy areas, upper beaches, rocky slopes, gravelly outwash fans, arroyos in tropical scrub, arid grasslands, desert scrub, pinyon-juniper woodlands Disturbed areas, gardens, road and railroad rights-of-way, stream beds
Elevation 0-2000 m (0-6600 ft) 0-1700 m [probably much higher in tropics] (0-5600 ft [probably much higher in tropics])
Distribution
from FNA
AL; AZ; CA; FL; LA; MD; NC; NM; NV; SC; TX; VA; Mexico; Central America; South America; West Indies; Eurasia; Africa; Australia
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; AZ; FL; GA; LA; MD; MO; MS; NC; NM; OK; SC; TN; TX; Mexico; Central America; South America; West Indies [Widely introduced throughout the tropics and warm-temperate regions]
[WildflowerSearch map]
[BONAP county map]
Discussion

Boerhavia coccinea is weedy and probably adventive along the Gulf and southern Atlantic coasts in North America; it can be expected in waste areas anywhere in that region. Worldwide, it probably also has a broader range than indicated, particularly on oceanic islands. The complex, which is in need of taxonomic clarification, is extremely variable with regard to robustness, pubescence, and fruit number in individual terminal inflorescences. In the New World, flowers are usually some shade of deep wine red, although populations of white-flowered or yellow-flowered plants are rarely found (R. Spellenberg 2000). In the Old World, pink-flowered plants are frequent (C. Whitehouse 1996).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Boerhavia erecta occasionally forms mixed populations with B. intermedia without apparent intergradation. Rarely, some specimens seem to combine features of either species, particularly with regard to inflorescence structure. This is especially so in Sonora, Mexico, and in parts of the Sonoran Desert in Arizona. The two species bloom simultaneously and are visited by small insects. Given the presumed close relationship and weedy habitats of each, hybridization seems possible. Usually, the two species can be distinguished by the differences in fruit length, the appearance of a crownlike apex of the nearly mature fruits of B. erecta (apex of ridges slightly expanded, apex of fruit slightly conic), and the more precisely constructed terminal umbels of B. intermedia. Both species, particularly B. intermedia, may produce entire inflorescences with branches terminating in single flowers. R. E. Woodson Jr. and H. J. Kidd (1961) suggested that B. erecta hybridizes with the perennial B. diffusa.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 4, p. 20. FNA vol. 4, p. 22.
Parent taxa Nyctaginaceae > Boerhavia Nyctaginaceae > Boerhavia
Sibling taxa
B. anisophylla, B. ciliata, B. coulteri, B. diffusa, B. erecta, B. gracillima, B. intermedia, B. linearifolia, B. megaptera, B. pterocarpa, B. purpurascens, B. spicata, B. torreyana, B. triquetra, B. wrightii
B. anisophylla, B. ciliata, B. coccinea, B. coulteri, B. diffusa, B. gracillima, B. intermedia, B. linearifolia, B. megaptera, B. pterocarpa, B. purpurascens, B. spicata, B. torreyana, B. triquetra, B. wrightii
Synonyms B. caribaea, B. viscosa
Name authority Miller: Gard. Dict. ed. 8, Boerhavia no. 4. (1768) Linnaeus: Sp. Pl. 1: 3. (1753)
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