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last chance rock cress

Habit Perennials; long-lived; sexual; caudex woody. Perennials; short-lived; sexual; caudex not woody.
Stems

usually 1 per caudex branch, arising from center of rosette near ground surface, 1–3 dm, densely pubescent proximally, trichomes short-stalked, 4–7-rayed, 0.1–0.5 mm, mixed proximally with simple and short- to long-stalked, 2- or 3-rayed ones, to 1.5 mm, moderately pubescent distally.

usually 1 per caudex branch, arising from center of rosette near ground surface, 0.7–2.2 dm, glabrous throughout.

Basal leaves

blade linear-oblanceolate, 1.5–3 mm wide, margins entire, ciliate proximally, trichomes (simple and 2- or 3-rayed), to 1.5 mm, surfaces moderately pubescent, trichomes short-stalked, 4–7-rayed, 0.3–0.6 mm.

blade oblanceolate, 1–2 mm wide, margins entire, ciliate near petiole base, trichomes (simple), to 0.4 mm, surfaces sparsely to densely pubescent, trichomes short-stalked, 4–8-rayed, 0.03–0.1 mm.

Cauline leaves

9–17, concealing stem proximally;

blade auricles absent, surfaces of distalmost leaves moderately pubescent.

6–11, sometimes concealing stem proximally;

blade auricles to 0.5 mm, surfaces of distalmost leaves glabrous.

Racemes

8–35-flowered, usually unbranched.

5–17-flowered, unbranched.

Flowers

pendent at anthesis;

sepals pubescent;

petals yellowish proximally and brick-red distally or, rarely, one color throughout, 9–10 × 0.8–1 mm, glabrous;

pollen ellipsoid.

erect at anthesis;

sepals glabrous;

petals white, 3–4 × 0.5–0.8 mm, glabrous;

pollen ellipsoid.

Fruiting pedicels

reflexed, straight, 1.5–2 mm, pubescent, trichomes appressed, branched.

erect, straight, 2–4(–7) mm, glabrous.

Fruits

(immature) reflexed, often appressed to rachis, not secund, straight, edges parallel, ca. 4 cm;

valves pubescent throughout;

style ca. 0.3 mm.

erect, usually appressed to rachis, not secund, straight, edges parallel, 2–3.5 cm × 2.2–2.5 mm;

valves glabrous;

ovules 60–96 per ovary;

style 0.05–0.2 mm.

Seeds

not seen.

sub-biseriate, 0.7–0.9 × 0.5–0.6 mm, not winged or minutely winged distally.

Boechera yorkii

Boechera williamsii

Phenology Flowering May. Flowering May–Jun.
Habitat Crevices and ledges of calcareous rock outcrops Gravelly soil in sagebrush-grassland communities
Elevation 2200-2400 m (7200-7900 ft) 2300-2800 m (7500-9200 ft)
Distribution
from FNA
CA
[BONAP county map]
from FNA
WY
[BONAP county map]
Discussion

Boechera yorkii is immediately recognizable by its yellowish to brick-red petals and extremely short (1.5–2 mm), reflexed fruiting pedicels. It is known only from Last Chance Mountains in Inyo County.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Boechera williamsii is distinguished from all other sexual species of the genus by its combination of relatively short (3–4 mm), white petals, minute, 4–8-rayed trichomes on basal leaves, erect, appressed fruits, and relatively small, wingless seeds. It is known only from Fremont and Sublette counties in west-central Wyoming. The taxon traditionally treated as Arabis (Boechera) williamsii var. saximontana is here recognized as an apomictic species of hybrid origin (see M. D. Windham and I. A. Al-Shehbaz 2007b for detailed comparison).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 412. FNA vol. 7, p. 411.
Parent taxa Brassicaceae > tribe Boechereae > Boechera Brassicaceae > tribe Boechereae > Boechera
Sibling taxa
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. xylopoda, B. yorkii
Synonyms Arabis williamsii
Name authority S. Boyd: Madroño 51: 387, figs. 1–3. (2004) (Rollins) Dorn: Vasc. Pl. Wyoming ed. 3, 376. (2001)
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