Boechera williamsii |
Boechera pratincola |
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meadow rock-cress |
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Habit | Perennials; short-lived; sexual; caudex not woody. | Perennials; usually short-lived; apomictic; caudex not woody. |
Stems | usually 1 per caudex branch, arising from center of rosette near ground surface, 0.7–2.2 dm, glabrous throughout. |
usually 1 per caudex branch, arising from center of rosette near ground surface, 2–6 dm, pubescent proximally, trichomes sessile, 2- or 3-rayed, 0.15–0.6 mm, glabrous distally. |
Basal leaves | blade oblanceolate, 1–2 mm wide, margins entire, ciliate near petiole base, trichomes (simple), to 0.4 mm, surfaces sparsely to densely pubescent, trichomes short-stalked, 4–8-rayed, 0.03–0.1 mm. |
blade oblanceolate, 1.5–5(–7) mm wide, margins usually entire, rarely denticulate, ciliate at petiole base, trichomes to 0.4 mm, surfaces pubescent, trichomes sessile, 2–4(–6)-rayed, 0.1–0.45 mm. |
Cauline leaves | 6–11, sometimes concealing stem proximally; blade auricles to 0.5 mm, surfaces of distalmost leaves glabrous. |
(7–)13–42, concealing stem proximally; blade auricles 1–4 mm, surfaces of distalmost leaves sparsely pubescent apically. |
Racemes | 5–17-flowered, unbranched. |
(7–)14–45-flowered, usually unbranched. |
Flowers | erect at anthesis; sepals glabrous; petals white, 3–4 × 0.5–0.8 mm, glabrous; pollen ellipsoid. |
ascending at anthesis; sepals glabrous; petals lavender, 6–10 × 1.5–2 mm, glabrous; pollen spheroid. |
Fruiting pedicels | erect, straight, 2–4(–7) mm, glabrous. |
erect, straight, 5–12 mm, glabrous. |
Fruits | erect, usually appressed to rachis, not secund, straight, edges parallel, 2–3.5 cm × 2.2–2.5 mm; valves glabrous; ovules 60–96 per ovary; style 0.05–0.2 mm. |
erect, appressed to rachis, not secund, straight, edges parallel, 4–6.5 cm × 1.5–2 mm; valves glabrous; ovules 60–140 per ovary; style 0.1–0.3 mm. |
Seeds | sub-biseriate, 0.7–0.9 × 0.5–0.6 mm, not winged or minutely winged distally. |
sub-biseriate, 1.4–2 × 1–1.4 mm; wing continuous, 0.1–0.35 mm wide. |
Boechera williamsii |
Boechera pratincola |
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Phenology | Flowering May–Jun. | Flowering Jun–Aug. |
Habitat | Gravelly soil in sagebrush-grassland communities | Rocky slopes and soil patches in open areas and along forest edges |
Elevation | 2300-2800 m (7500-9200 ft) | 1900-3200 m (6200-10500 ft) |
Distribution |
WY |
CA; ID; NV; OR
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Discussion | Boechera williamsii is distinguished from all other sexual species of the genus by its combination of relatively short (3–4 mm), white petals, minute, 4–8-rayed trichomes on basal leaves, erect, appressed fruits, and relatively small, wingless seeds. It is known only from Fremont and Sublette counties in west-central Wyoming. The taxon traditionally treated as Arabis (Boechera) williamsii var. saximontana is here recognized as an apomictic species of hybrid origin (see M. D. Windham and I. A. Al-Shehbaz 2007b for detailed comparison). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Morphological evidence suggests that Boechera pratincola is an apomictic species that arose through hybridization between B. paupercula and B. stricta (see M. D. Windham and I. A. Al-Shehbaz 2007b for detailed comparison). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 411. | FNA vol. 7, p. 399. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Arabis williamsii | Arabis pratincola, Arabis drummondii var. pratincola, Arabis nemophila |
Name authority | (Rollins) Dorn: Vasc. Pl. Wyoming ed. 3, 376. (2001) | (Greene) Windham & Al-Shehbaz: Harvard Pap. Bot. 12: 245. (2007) |
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