Boechera williamsii |
Boechera hastatula |
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Hells Canyon rock-cress |
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Habit | Perennials; short-lived; sexual; caudex not woody. | Perennials; short- to long-lived; apomictic or sexual; caudex often woody. |
Stems | usually 1 per caudex branch, arising from center of rosette near ground surface, 0.7–2.2 dm, glabrous throughout. |
1–4 per caudex branch, arising from margin of rosette near ground surface, 1–3 dm, glabrous throughout. |
Basal leaves | blade oblanceolate, 1–2 mm wide, margins entire, ciliate near petiole base, trichomes (simple), to 0.4 mm, surfaces sparsely to densely pubescent, trichomes short-stalked, 4–8-rayed, 0.03–0.1 mm. |
blade linear-oblanceolate to oblanceolate, 1–3(–5) mm wide, margins entire or minutely toothed, rarely ciliate near petiole base, trichomes (simple), to 0.8 mm, surfaces pubescent or glabrous, trichomes short-stalked, 3–6-rayed, 0.07–0.2 mm. |
Cauline leaves | 6–11, sometimes concealing stem proximally; blade auricles to 0.5 mm, surfaces of distalmost leaves glabrous. |
4–15, not concealing stem; blade auricles 0.5–2.5 mm, surfaces of distalmost leaves glabrous. |
Racemes | 5–17-flowered, unbranched. |
12–40-flowered, usually unbranched. |
Flowers | erect at anthesis; sepals glabrous; petals white, 3–4 × 0.5–0.8 mm, glabrous; pollen ellipsoid. |
divaricate to pendent at anthesis; sepals pubescent; petals lavender, 5–6 × ca. 1 mm, glabrous; pollen ellipsoid or spheroid. |
Fruiting pedicels | erect, straight, 2–4(–7) mm, glabrous. |
reflexed, abruptly recurved near base, 4–8 mm, glabrous. |
Fruits | erect, usually appressed to rachis, not secund, straight, edges parallel, 2–3.5 cm × 2.2–2.5 mm; valves glabrous; ovules 60–96 per ovary; style 0.05–0.2 mm. |
reflexed to closely pendent, usually appressed to rachis, often secund, straight, edges parallel, 0.3–0.42 cm × 1–1.5 mm; valves glabrous; ovules 46–60 per ovary. |
Seeds | sub-biseriate, 0.7–0.9 × 0.5–0.6 mm, not winged or minutely winged distally. |
uniseriate, 1–1.3 × 0.8–1 mm; wing continuous, 0.05–0.1 mm wide. |
Boechera williamsii |
Boechera hastatula |
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Phenology | Flowering May–Jun. | Flowering Jun–Jul. |
Habitat | Gravelly soil in sagebrush-grassland communities | Igneous rock outcrops |
Elevation | 2300-2800 m (7500-9200 ft) | 1900-2100 m (6200-6900 ft) |
Distribution |
WY |
OR |
Discussion | Boechera williamsii is distinguished from all other sexual species of the genus by its combination of relatively short (3–4 mm), white petals, minute, 4–8-rayed trichomes on basal leaves, erect, appressed fruits, and relatively small, wingless seeds. It is known only from Fremont and Sublette counties in west-central Wyoming. The taxon traditionally treated as Arabis (Boechera) williamsii var. saximontana is here recognized as an apomictic species of hybrid origin (see M. D. Windham and I. A. Al-Shehbaz 2007b for detailed comparison). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Both sexual and apomictic collections of Boechera hastatula have been identified; further study is needed to determine whether they truly are conspecific. The species is known only from Baker and Wallowa counties. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 411. | FNA vol. 7, p. 383. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Arabis williamsii | Arabis hastatula |
Name authority | (Rollins) Dorn: Vasc. Pl. Wyoming ed. 3, 376. (2001) | (Greene) Al-Shehbaz: Novon 13: 386. (2003) |
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