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Hells Canyon rock-cress

Habit Perennials; short-lived; sexual; caudex not woody. Perennials; short- to long-lived; apomictic or sexual; caudex often woody.
Stems

usually 1 per caudex branch, arising from center of rosette near ground surface, 0.7–2.2 dm, glabrous throughout.

1–4 per caudex branch, arising from margin of rosette near ground surface, 1–3 dm, glabrous throughout.

Basal leaves

blade oblanceolate, 1–2 mm wide, margins entire, ciliate near petiole base, trichomes (simple), to 0.4 mm, surfaces sparsely to densely pubescent, trichomes short-stalked, 4–8-rayed, 0.03–0.1 mm.

blade linear-oblanceolate to oblanceolate, 1–3(–5) mm wide, margins entire or minutely toothed, rarely ciliate near petiole base, trichomes (simple), to 0.8 mm, surfaces pubescent or glabrous, trichomes short-stalked, 3–6-rayed, 0.07–0.2 mm.

Cauline leaves

6–11, sometimes concealing stem proximally;

blade auricles to 0.5 mm, surfaces of distalmost leaves glabrous.

4–15, not concealing stem;

blade auricles 0.5–2.5 mm, surfaces of distalmost leaves glabrous.

Racemes

5–17-flowered, unbranched.

12–40-flowered, usually unbranched.

Flowers

erect at anthesis;

sepals glabrous;

petals white, 3–4 × 0.5–0.8 mm, glabrous;

pollen ellipsoid.

divaricate to pendent at anthesis;

sepals pubescent;

petals lavender, 5–6 × ca. 1 mm, glabrous;

pollen ellipsoid or spheroid.

Fruiting pedicels

erect, straight, 2–4(–7) mm, glabrous.

reflexed, abruptly recurved near base, 4–8 mm, glabrous.

Fruits

erect, usually appressed to rachis, not secund, straight, edges parallel, 2–3.5 cm × 2.2–2.5 mm;

valves glabrous;

ovules 60–96 per ovary;

style 0.05–0.2 mm.

reflexed to closely pendent, usually appressed to rachis, often secund, straight, edges parallel, 0.3–0.42 cm × 1–1.5 mm;

valves glabrous;

ovules 46–60 per ovary.

Seeds

sub-biseriate, 0.7–0.9 × 0.5–0.6 mm, not winged or minutely winged distally.

uniseriate, 1–1.3 × 0.8–1 mm;

wing continuous, 0.05–0.1 mm wide.

Boechera williamsii

Boechera hastatula

Phenology Flowering May–Jun. Flowering Jun–Jul.
Habitat Gravelly soil in sagebrush-grassland communities Igneous rock outcrops
Elevation 2300-2800 m (7500-9200 ft) 1900-2100 m (6200-6900 ft)
Distribution
from FNA
WY
[BONAP county map]
from FNA
OR
[BONAP county map]
Discussion

Boechera williamsii is distinguished from all other sexual species of the genus by its combination of relatively short (3–4 mm), white petals, minute, 4–8-rayed trichomes on basal leaves, erect, appressed fruits, and relatively small, wingless seeds. It is known only from Fremont and Sublette counties in west-central Wyoming. The taxon traditionally treated as Arabis (Boechera) williamsii var. saximontana is here recognized as an apomictic species of hybrid origin (see M. D. Windham and I. A. Al-Shehbaz 2007b for detailed comparison).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Both sexual and apomictic collections of Boechera hastatula have been identified; further study is needed to determine whether they truly are conspecific. The species is known only from Baker and Wallowa counties.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 411. FNA vol. 7, p. 383.
Parent taxa Brassicaceae > tribe Boechereae > Boechera Brassicaceae > tribe Boechereae > Boechera
Sibling taxa
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. xylopoda, B. yorkii
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hoffmannii, B. holboellii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
Synonyms Arabis williamsii Arabis hastatula
Name authority (Rollins) Dorn: Vasc. Pl. Wyoming ed. 3, 376. (2001) (Greene) Al-Shehbaz: Novon 13: 386. (2003)
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