Boechera villosa |
Boechera johnstonii |
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Johnston's rockcress |
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Habit | Perennials; short-lived; sexual; caudex not woody. | Perennials; long-lived; (cespitose); sexual; caudex woody. |
Stems | 1–3 per caudex branch, arising laterally proximal to sterile shoots near ground surface, ca. 2.5 dm, densely pubescent proximally, trichomes simple and 2-rayed, 0.25–0.7 mm, glabrescent distally. |
usually 1 per caudex branch, arising from center of rosette near ground surface, 0.5–2 dm, densely pubescent proximally, trichomes short-stalked, 4–10-rayed, 0.07–0.15 mm, sparsely to densely pubescent distally. |
Basal leaves | blade oblanceolate to obovate, 2.5–5 mm wide, margins entire, ciliate along petiole, trichomes (simple), to 1 mm, surfaces densely pubescent, trichomes simple and short-stalked, 2- or 3-rayed, 0.25–0.5 mm. |
blade narrowly oblanceolate, 1.5–4 mm wide, margins entire, not ciliate, surfaces densely pubescent, trichomes short-stalked, 6–14-rayed, 0.07–0.15 mm. |
Cauline leaves | 4–6, not concealing stem; blade auricles ca. 1 mm, surfaces of distalmost leaves sparsely pubescent. |
4–10, often concealing stem proximally; blade auricles absent, surfaces of distalmost leaves densely pubescent. |
Racemes | 6–10-flowered, unbranched. |
10–18-flowered, unbranched. |
Flowers | ascending at anthesis; sepals pubescent; petals lavender, 4–5 × 1–1.5 mm, glabrous; pollen ellipsoid. |
ascending at anthesis; sepals pubescent; petals purple, 9–14 × 2–4 mm, glabrous; pollen ellipsoid. |
Fruiting pedicels | ascending, straight, 6–10 mm, glabrous or sparsely pubescent, trichomes subappressed, branched. |
divaricate-ascending, straight, 5–14 mm, pubescent, trichomes appressed, branched. |
Fruits | divaricate-ascending, not appressed to rachis, not secund, slightly curved, edges parallel, 4–5 cm × ca. 1 mm; valves glabrous; ovules ca. 64 per ovary; style ca. 0.2 mm. |
divaricate-ascending, not appressed to rachis, not secund, straight, edges parallel, 4–6 cm × 2.5–4 mm; valves glabrous; ovules 26–34 per ovary; style (0.7–)1–2 mm. |
Seeds | uniseriate, (none mature). |
uniseriate, 1.9–2.7 × 1.5–2.2 mm; wing continuous, 0.3–0.7 mm wide. |
Boechera villosa |
Boechera johnstonii |
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Phenology | Flowering May. | Flowering Feb–Mar. |
Habitat | Basalt outcrop in pinyon-juniper woodlands | Rocky areas and gravelly soil in chaparral and oak-pine savannas |
Elevation | ca. 2100 m (ca. 6900 ft) | 1300-1700 m (4300-5600 ft) |
Distribution |
NM |
CA |
Discussion | Boechera villosa is known only from the type collection from the Rio Grande Gorge in Taos County. The holotype was originally identified as Arabis (Boechera) perennans but clearly is more closely related to B. pallidifolia (see M. D. Windham and I. A. Al-Shehbaz 2006 for detailed comparison). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Boechera johnstonii and B. hirshbergiae are too similar to be considered distinct species. Even with the expanded circumscription, this distinctive sexual species is known only from the area of Cuyamaca Lake (San Diego County) and the San Jacinto Mountains (Riverside County). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 411. | FNA vol. 7, p. 386. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Arabis johnstonii, Arabis hirshbergiae, B. hirshbergiae | |
Name authority | Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 86. (2006) | (Munz) Al-Shehbaz: Novon 13: 386. (2003) |
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