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Coville's rock-cress

Habit Perennials; short-lived; sexual; caudex not woody. Perennials; long-lived; apomictic; caudex woody.
Stems

1–3 per caudex branch, arising laterally proximal to sterile shoots near ground surface, ca. 2.5 dm, densely pubescent proximally, trichomes simple and 2-rayed, 0.25–0.7 mm, glabrescent distally.

usually 1 per caudex branch, arising from center of rosette near ground surface, 0.5–2.5 dm, glabrous throughout.

Basal leaves

blade oblanceolate to obovate, 2.5–5 mm wide, margins entire, ciliate along petiole, trichomes (simple), to 1 mm, surfaces densely pubescent, trichomes simple and short-stalked, 2- or 3-rayed, 0.25–0.5 mm.

blade oblanceolate, 1.5–5 mm wide, margins entire, ciliate proximally, trichomes (simple), to 0.4 mm, and distally, trichomes short-stalked, 2 or 3 (or 4)-rayed, 0.15–0.4 mm, surfaces glabrous.

Cauline leaves

4–6, not concealing stem;

blade auricles ca. 1 mm, surfaces of distalmost leaves sparsely pubescent.

2–7, not concealing stem;

blade with auricles 0.2–1 mm, auricles rarely absent, surfaces of distalmost leaves glabrous.

Racemes

6–10-flowered, unbranched.

2–9-flowered, unbranched.

Flowers

ascending at anthesis;

sepals pubescent;

petals lavender, 4–5 × 1–1.5 mm, glabrous;

pollen ellipsoid.

ascending at anthesis;

sepals glabrous;

petals lavender to purplish, 5–6 × 1–2 mm, glabrous;

pollen spheroid.

Fruiting pedicels

ascending, straight, 6–10 mm, glabrous or sparsely pubescent, trichomes subappressed, branched.

ascending to suberect, straight, 6–20 mm, glabrous.

Fruits

divaricate-ascending, not appressed to rachis, not secund, slightly curved, edges parallel, 4–5 cm × ca. 1 mm;

valves glabrous;

ovules ca. 64 per ovary;

style ca. 0.2 mm.

ascending to suberect, rarely appressed to rachis, not secund, straight, edges slightly undulate (rarely parallel), 3.5–5 cm × 3–5 mm;

valves glabrous;

ovules 18–30 per ovary;

style 0.2–1 mm.

Seeds

uniseriate, (none mature).

uniseriate, 4–5 × 3–4 mm;

wing continuous, 1–1.5 mm wide.

Boechera villosa

Boechera covillei

Phenology Flowering May. Flowering Jul–Aug.
Habitat Basalt outcrop in pinyon-juniper woodlands Rocky slopes in alpine meadows and open coniferous forests
Elevation ca. 2100 m (ca. 6900 ft) 2200-3500 m (7200-11500 ft)
Distribution
from FNA
NM
[BONAP county map]
from FNA
CA; NV; OR
[BONAP county map]
Discussion

Boechera villosa is known only from the type collection from the Rio Grande Gorge in Taos County. The holotype was originally identified as Arabis (Boechera) perennans but clearly is more closely related to B. pallidifolia (see M. D. Windham and I. A. Al-Shehbaz 2006 for detailed comparison).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Morphological evidence suggests that Boechera covillei is an apomictic species that arose through hybridization between B. howellii and B. lyallii (see M. D. Windham and I. A. Al-Shehbaz 2007b for detailed comparison).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 411. FNA vol. 7, p. 371.
Parent taxa Brassicaceae > tribe Boechereae > Boechera Brassicaceae > tribe Boechereae > Boechera
Sibling taxa
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. williamsii, B. xylopoda, B. yorkii
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
Synonyms Arabis covillei, Arabis leibergii
Name authority Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 86. (2006) (Greene) Windham & Al-Shehbaz: Harvard Pap. Bot. 12: 237. (2007)
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