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smooth rockcress

Habit Perennials; long-lived; reproductive mode unknown; caudex woody. Biennials; short-lived; sexual; caudex not evident.
Stems

1 per caudex branch, arising from center of rosette near ground surface, ca. 1 dm, densely pubescent proximally, trichomes stalked, (2 or) 3–6-rayed, 0.2–0.5 mm, glabrate distally.

1 per plant, arising from center of rosette near ground surface, (1.5–)3–11 dm, glabrous throughout.

Basal leaves

blade oblanceolate, 4–6 mm wide, margins entire, not ciliate, surfaces moderately pubescent, trichomes long-stalked, 3–6 (or 7)-rayed, 0.2–0.5 mm.

blade obovate to oblanceolate, (4–)10–40 mm wide, margins serrate or dentate, ciliate-mucronate on teeth, trichomes often minute, surfaces glabrous or sparsely pubescent, trichomes simple, 0.1–0.6 mm.

Cauline leaves

2–5, concealing stem proximally;

blade auricles absent, surfaces of distalmost leaves pubescent.

7–15, often concealing stem proximally;

blade auricles 3–12(–17) mm, surfaces of distalmost leaves glabrous.

Racemes

3- or 4-flowered, unbranched.

16–45-flowered, sometimes branched.

Flowers

not seen.

ascending at anthesis;

sepals glabrous;

petals white, 3–5 × 1–1.5 mm, glabrous;

pollen ellipsoid.

Fruiting pedicels

erect-ascending, straight, 4–5 mm, glabrous or sparsely pubescent, trichomes appressed, branched.

suberect to divaricate-ascending, straight to slightly curved, 5–23 mm, glabrous.

Fruits

erect-ascending, sometimes appressed to rachis, not secund, straight, edges parallel, 3–4 cm × ca. 5 mm;

valves glabrous;

ovules ca. 16 per ovary;

style 0.5–0.7 mm.

divaricate-ascending, not appressed to rachis, rarely somewhat secund, curved, edges parallel, (4–)6–11.7 cm × 1–2 mm;

valves glabrous;

ovules 50–80 per ovary;

style 0.1–0.7(–1) mm.

Seeds

uniseriate, 5.5–6.5 × 4–4.5 mm;

wing continuous, 1–2 mm wide.

uniseriate, 1.2–2.2 × 0.8–1.4 mm;

wing continuous, 0.1–0.3 mm wide distally.

2n

= 14.

Boechera ultra-alsa

Boechera laevigata

Phenology Flowering Jun–Jul. Flowering Mar–May.
Habitat Rocky soil Rocky bluffs, cedar glades, wooded hillsides, floodplains
Elevation ca. 1800 m (ca. 5900 ft) 100-500 m (300-1600 ft)
Distribution
from FNA
CA
[BONAP county map]
from FNA
AL; AR; CT; DC; DE; GA; IA; IL; IN; KY; MA; MD; MI; MO; NC; NH; NJ; NY; OH; OK; PA; SC; TN; VA; VT; WI; WV; ON; QC
Discussion

Boechera ultra-alsa is known only from the holotype, from Snow Mountain in Lake County. Originally identified as Arabis (Boechera) platysperma, this specimen differs from that taxon by having petioles without obvious cilia, basal leaves with long-stalked (0.1–0.2 versus less than 0.1 mm) trichomes, and fruits edges parallel (not undulate or constricted between the seeds).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

The taxon sometimes treated as Arabis laevigata var. missouriensis is here recognized as a separate species (Boechera missouriensis) based on its significantly longer petals (5–10 versus 3–5 mm) and distinctive, lyrate-pinnatifid basal leaves that persist well beyond anthesis.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 410. FNA vol. 7, p. 386.
Parent taxa Brassicaceae > tribe Boechereae > Boechera Brassicaceae > tribe Boechereae > Boechera
Sibling taxa
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. villosa, B. williamsii, B. xylopoda, B. yorkii
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
Synonyms Turritis laevigata, Arabis hastata, Arabis heterophylla, Arabis laevigata, Arabis lyrifolia
Name authority Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 86. (2006) (Muhlenberg ex Willdenow) Al-Shehbaz: Novon 13: 386. (2003)
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