Boechera ultra-alsa |
Boechera inyoensis |
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Inyo rockcress |
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Habit | Perennials; long-lived; reproductive mode unknown; caudex woody. | Perennials; short-lived; apomictic; caudex usually not woody. |
Stems | 1 per caudex branch, arising from center of rosette near ground surface, ca. 1 dm, densely pubescent proximally, trichomes stalked, (2 or) 3–6-rayed, 0.2–0.5 mm, glabrate distally. |
usually 1 per caudex, arising from center of rosette near ground surface, (1–)2.5–6.5 dm, densely pubescent proximally, trichomes short-stalked, 7–12-rayed, 0.1–0.2 mm, densely to sparsely pubescent distally. |
Basal leaves | blade oblanceolate, 4–6 mm wide, margins entire, not ciliate, surfaces moderately pubescent, trichomes long-stalked, 3–6 (or 7)-rayed, 0.2–0.5 mm. |
blade oblanceolate, 1–4(–8) mm wide, margins entire, ciliate along petiole and proximally, trichomes (simple), to 1 mm, surfaces densely pubescent, trichomes short-stalked, 3–10-rayed, 0.2–0.7 mm. |
Cauline leaves | 2–5, concealing stem proximally; blade auricles absent, surfaces of distalmost leaves pubescent. |
(7–)12–35, concealing stem proximally; blade auricles 0.5–2 mm, surfaces of distalmost leaves densely pubescent. |
Racemes | 3- or 4-flowered, unbranched. |
10–65-flowered, usually unbranched. |
Flowers | not seen. |
ascending at anthesis; sepals pubescent; petals lavender to purplish, 5–8 × 1.2–2 mm, glabrous; pollen spheroid. |
Fruiting pedicels | erect-ascending, straight, 4–5 mm, glabrous or sparsely pubescent, trichomes appressed, branched. |
divaricate-ascending to horizontal, straight or slightly curved downward, 5–15 mm, sparsely pubescent, trichomes appressed, branched. |
Fruits | erect-ascending, sometimes appressed to rachis, not secund, straight, edges parallel, 3–4 cm × ca. 5 mm; valves glabrous; ovules ca. 16 per ovary; style 0.5–0.7 mm. |
divaricate-ascending to widely pendent, not appressed to rachis, not secund, usually curved, rarely straight, edges parallel, 3.7–6.5 cm × 1.5–2.2 mm; valves glabrous; ovules 74–134 per ovary; style 0.05–0.2 mm. |
Seeds | uniseriate, 5.5–6.5 × 4–4.5 mm; wing continuous, 1–2 mm wide. |
usually sub-biseriate, rarely uniseriate, 1.7–2 × 1–1.5 mm; wing continuous, 0.1–0.2 mm wide. |
2n | = 21. |
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Boechera ultra-alsa |
Boechera inyoensis |
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Phenology | Flowering Jun–Jul. | Flowering Apr–Jun. |
Habitat | Rocky soil | Limestone and volcanic rock outcrops and clay soils in desert scrub and pinyon-juniper woodlands |
Elevation | ca. 1800 m (ca. 5900 ft) | 1400-2400 m (4600-7900 ft) |
Distribution |
CA |
CA; NV; UT
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Discussion | Boechera ultra-alsa is known only from the holotype, from Snow Mountain in Lake County. Originally identified as Arabis (Boechera) platysperma, this specimen differs from that taxon by having petioles without obvious cilia, basal leaves with long-stalked (0.1–0.2 versus less than 0.1 mm) trichomes, and fruits edges parallel (not undulate or constricted between the seeds). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Some plants here assigned to Boechera inyoensis were treated by R. C. Rollins (1993) and N. H. Holmgren (2005b) as Arabis (or B.) beckwithii. Closer examination of the type specimens of A. beckwithii reveals that they are conspecific with B. puberula, a sexual diploid widespread in the Great Basin of western North America. Boechera inyoensis is an apomictic triploid that clearly contains at least one genome derived from B. shockleyi (M. D. Windham and I. A. Al-Shehbaz 2007b); the other parent has not been determined. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 410. | FNA vol. 7, p. 385. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Arabis inyoensis, Arabis holboellii var. derensis, B. selbyi var. inyoensis | |
Name authority | Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 86. (2006) | (Rollins) Al-Shehbaz: Novon 13: 386. (2003) |
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