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Tulare rockcress

Trinity Mountain rockcress

Habit Biennials or perennials; short-lived; apomictic; caudex present or absent. Perennials; long-lived; apomictic; caudex woody.
Stems

usually 1 per caudex branch, arising from center of rosette near ground surface, 2–7 dm, sparsely to densely pubescent proximally, trichomes subsessile, sub-malpighiaceous, 0.3–0.6 mm, glabrous distally.

usually 1 per caudex branch, arising from center of rosette, slightly elevated on woody base or from near ground surface, 2–6 dm, pubescent proximally, trichomes usually subsessile, 2- or 3-rayed, rarely simple, 0.25–0.4 mm, glabrous distally.

Basal leaves

blade oblanceolate, 3–7 mm wide, margins entire, ciliate proximally, trichomes (simple), to 0.8 mm, surfaces sparsely to densely pubescent, trichomes subsessile, 2–5-rayed, 0.2–0.55 mm.

blade oblanceolate, 2–6 mm wide, margins entire, ciliate proximally, trichomes to 0.7 mm, surfaces sparsely pubescent, trichomes subsessile to short-stalked, 2–5-rayed, 0.1–0.4 mm.

Cauline leaves

7–17, often concealing stem proximally;

blade auricles 2–5 mm, surfaces of distalmost leaves sparsely pubescent or glabrous.

5–10, rarely concealing stem proximally;

blade auricles 0.5–2 mm, surfaces of distalmost leaves glabrous.

Racemes

19–39-flowered, usually unbranched.

5–16-flowered, usually unbranched.

Flowers

divaricate to pendent at anthesis;

sepals pubescent;

petals white to pale lavender, 6–7 × 1.2–2 mm, glabrous;

pollen spheroid.

ascending at anthesis;

sepals glabrous;

petals purple, 6–8 × 1.5–2 mm, glabrous;

pollen spheroid.

Fruiting pedicels

reflexed, recurved proximally, 5–13 mm, glabrous.

divaricate-ascending, straight, 4–10 mm, glabrous.

Fruits

reflexed, rarely appressed to rachis, not secund, straight, edges parallel, 4–7(–8.5) cm × 2–2.3 mm;

valves glabrous;

ovules 88–104 per ovary;

style 0.3–0.7 mm.

divaricate-ascending, not appressed to rachis, not secund, straight, edges somewhat undulate (not parallel), 4–6.5(–7.6) cm × 2.5–3.5 mm;

valves glabrous;

ovules 24–54 per ovary;

style 0.2–0.8 mm.

Seeds

sub-biseriate, 2–2.5 × 1.1–1.5 mm;

wing continuous, 0.15–0.25 mm wide.

uniseriate, 2.5–3.2 × 1.8–2.5 mm;

wing distal, 0.3–1 mm wide.

Boechera tularensis

Boechera rigidissima

Phenology Flowering Jun–Jul. Flowering Jul–Aug.
Habitat Rocky slopes in montane and subalpine habitats Rocky places in open conifer forests
Elevation 2400-3200 m (7900-10500 ft) 1800-2000 m (5900-6600 ft)
Distribution
from FNA
CA
[BONAP county map]
from FNA
CA; NV
[BONAP county map]
Discussion

Morphological evidence suggests that Boechera tularensis is an apomictic species that contains three different genomes, one each from B. rectissima, B. retrofracta, and B. stricta. It is most often confused with B. pinetorum and B. retrofracta, but is easily distinguishable from both (see M. D. Windham and I. A. Al-Shehbaz 2007b for detailed comparison). It is known from the southern Sierra Nevada (Fresno and Tulare counties).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Plants assigned to Boechera rigidissima appear to represent a series of apomictic hybrids, possibly involving B. stricta and B. suffrutescens. The two varieties recognized by Rollins are well isolated geographically (in Trinity County, California, and Washoe County, Nevada, respectively) and show enough morphological divergence to suggest that they represent distinct origins, if not different combinations of parental genomes.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 410. FNA vol. 7, p. 403.
Parent taxa Brassicaceae > tribe Boechereae > Boechera Brassicaceae > tribe Boechereae > Boechera
Sibling taxa
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. repanda, B. retrofracta, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
Synonyms Arabis rigidissima, Arabis rigidissima var. demota
Name authority Windham & Al-Shehbaz: Harvard Pap. Bot. 12: 249. (2007) (Rollins) Al-Shehbaz: Novon 13: 389. (2003)
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