Boechera tularensis |
Boechera fernaldiana |
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Tulare rockcress |
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Habit | Biennials or perennials; short-lived; apomictic; caudex present or absent. | Perennials; long-lived; (cespitose); sexual; caudex somewhat woody (sometimes with persistent, crowded leaf bases). | ||||
Stems | usually 1 per caudex branch, arising from center of rosette near ground surface, 2–7 dm, sparsely to densely pubescent proximally, trichomes subsessile, sub-malpighiaceous, 0.3–0.6 mm, glabrous distally. |
usually 1 per caudex branch, arising from center of rosette, near ground surface or slightly elevated on woody base, 1–3.8 dm, densely pubescent proximally, trichomes short-stalked, 4–8-rayed, 0.04–0.1 mm, sometimes mixed with stalked, 2-rayed ones, to 0.5 mm, usually sparsely pubescent distally. |
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Basal leaves | blade oblanceolate, 3–7 mm wide, margins entire, ciliate proximally, trichomes (simple), to 0.8 mm, surfaces sparsely to densely pubescent, trichomes subsessile, 2–5-rayed, 0.2–0.55 mm. |
blade narrowly oblanceolate, 1–4 mm wide, margins entire, ciliate along petiole, trichomes (simple), to 1 mm, surfaces densely pubescent, trichomes short-stalked, 4–8-rayed, 0.04–0.1 mm. |
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Cauline leaves | 7–17, often concealing stem proximally; blade auricles 2–5 mm, surfaces of distalmost leaves sparsely pubescent or glabrous. |
5–10, not concealing stem; blade auricles 0.3–2 mm, surfaces of distalmost leaves usually sparsely pubescent. |
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Racemes | 19–39-flowered, usually unbranched. |
10–20-flowered, usually unbranched. |
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Flowers | divaricate to pendent at anthesis; sepals pubescent; petals white to pale lavender, 6–7 × 1.2–2 mm, glabrous; pollen spheroid. |
ascending at anthesis; sepals (purplish or greenish) glabrous or sparsely or moderately pubescent; petals white or purple to lavender, 7–12 × 1.5–4 mm, glabrous; pollen ellipsoid. |
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Fruiting pedicels | reflexed, recurved proximally, 5–13 mm, glabrous. |
divaricate-ascending, straight, 5–20 mm, sparsely pubescent, trichomes appressed, branched. |
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Fruits | reflexed, rarely appressed to rachis, not secund, straight, edges parallel, 4–7(–8.5) cm × 2–2.3 mm; valves glabrous; ovules 88–104 per ovary; style 0.3–0.7 mm. |
divaricate-ascending, not appressed to rachis, not secund, straight or curved, edges parallel, 3.5–6.5(–7.5) cm × 1–1.6 mm; valves glabrous; ovules 30–72 per ovary; style 0.1–1 mm. |
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Seeds | sub-biseriate, 2–2.5 × 1.1–1.5 mm; wing continuous, 0.15–0.25 mm wide. |
uniseriate, 1–1.2 × 0.8–1 mm; wing continuous, ca. 0.1 mm wide. |
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Boechera tularensis |
Boechera fernaldiana |
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Phenology | Flowering Jun–Jul. | |||||
Habitat | Rocky slopes in montane and subalpine habitats | |||||
Elevation | 2400-3200 m (7900-10500 ft) | |||||
Distribution |
CA |
CO; NV; UT |
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Discussion | Morphological evidence suggests that Boechera tularensis is an apomictic species that contains three different genomes, one each from B. rectissima, B. retrofracta, and B. stricta. It is most often confused with B. pinetorum and B. retrofracta, but is easily distinguishable from both (see M. D. Windham and I. A. Al-Shehbaz 2007b for detailed comparison). It is known from the southern Sierra Nevada (Fresno and Tulare counties). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Subspecies 2 (2 in the flora). Taxa treated here as subspecies of Boechera fernaldiana share many morphological traits, and most recent authors (R. C. Rollins 1993; N. H. Holmgren 2005b) did not recognize them as distinct. They are consistently separated by the characters listed below, show some degree of molecular divergence (C. D. Bailey et al., unpubl.), and their ranges are separated by ca. 500 km. They clearly represent genetically isolated population systems that warrant taxonomic recognition. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 410. | FNA vol. 7, p. 378. | ||||
Parent taxa | ||||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Arabis fernaldiana, Arabis canescens var. stylosa, Arabis fernaldiana var. stylosa | |||||
Name authority | Windham & Al-Shehbaz: Harvard Pap. Bot. 12: 249. (2007) | (Rollins) W. A. Weber: Phytologia 51: 370. (1982) | ||||
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