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Tulare rockcress

Elko rockcress

Habit Biennials or perennials; short-lived; apomictic; caudex present or absent. Perennials; usually long-lived; apomictic; caudex often woody.
Stems

usually 1 per caudex branch, arising from center of rosette near ground surface, 2–7 dm, sparsely to densely pubescent proximally, trichomes subsessile, sub-malpighiaceous, 0.3–0.6 mm, glabrous distally.

1–5 per caudex branch, arising from center or margin of rosette near ground surface, 1–3 dm, sparsely pubescent proximally, trichomes subsessile, 2–5-rayed, 0.1–0.4 mm, glabrous or sparsely pubescent distally.

Basal leaves

blade oblanceolate, 3–7 mm wide, margins entire, ciliate proximally, trichomes (simple), to 0.8 mm, surfaces sparsely to densely pubescent, trichomes subsessile, 2–5-rayed, 0.2–0.55 mm.

blade narrowly oblanceolate, 2–6 mm wide, margins entire, ciliate at petiole base, trichomes (simple), to 0.4 mm, surfaces moderately pubescent, trichomes sessile or subsessile, 2–5-rayed, 0.1–0.4 mm.

Cauline leaves

7–17, often concealing stem proximally;

blade auricles 2–5 mm, surfaces of distalmost leaves sparsely pubescent or glabrous.

3–6, not concealing stem;

blade auricles absent or 0.2–0.5 mm, surfaces of distalmost leaves glabrous, or sparsely pubescent along margins.

Racemes

19–39-flowered, usually unbranched.

4–11-flowered, unbranched.

Flowers

divaricate to pendent at anthesis;

sepals pubescent;

petals white to pale lavender, 6–7 × 1.2–2 mm, glabrous;

pollen spheroid.

erect to ascending at anthesis;

sepals glabrous;

petals usually lavender, rarely whitish, 4–7 × 1.2–2 mm, glabrous;

pollen spheroid.

Fruiting pedicels

reflexed, recurved proximally, 5–13 mm, glabrous.

erect to ascending, straight, 4–8 mm, usually glabrous, rarely sparsely pubescent, trichomes appressed, branched.

Fruits

reflexed, rarely appressed to rachis, not secund, straight, edges parallel, 4–7(–8.5) cm × 2–2.3 mm;

valves glabrous;

ovules 88–104 per ovary;

style 0.3–0.7 mm.

erect to ascending, often appressed to rachis, not secund, straight, edges often slightly undulate (not parallel), 3–7 cm × 2–3 mm;

valves glabrous;

ovules 26–42 per ovary;

style 0.05–0.3 mm.

Seeds

sub-biseriate, 2–2.5 × 1.1–1.5 mm;

wing continuous, 0.15–0.25 mm wide.

uniseriate, 2.5–3.5 × 1.8–2.5 mm;

wing continuous, 0.4–0.9 mm wide.

Boechera tularensis

Boechera elkoensis

Phenology Flowering Jun–Jul. Flowering Jul.
Habitat Rocky slopes in montane and subalpine habitats Gravelly soil among rocks in open forests and subalpine meadows
Elevation 2400-3200 m (7900-10500 ft) 2000-3200 m (6600-10500 ft)
Distribution
from FNA
CA
[BONAP county map]
from FNA
CA; NV
[BONAP county map]
Discussion

Morphological evidence suggests that Boechera tularensis is an apomictic species that contains three different genomes, one each from B. rectissima, B. retrofracta, and B. stricta. It is most often confused with B. pinetorum and B. retrofracta, but is easily distinguishable from both (see M. D. Windham and I. A. Al-Shehbaz 2007b for detailed comparison). It is known from the southern Sierra Nevada (Fresno and Tulare counties).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Morphological evidence suggests that Boechera elkoensis is an apomictic species that arose through hybridization between B. platysperma and B. stricta (see M. D. Windham and I. A. Al-Shehbaz 2007 for detailed comparison).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 410. FNA vol. 7, p. 375.
Parent taxa Brassicaceae > tribe Boechereae > Boechera Brassicaceae > tribe Boechereae > Boechera
Sibling taxa
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
Name authority Windham & Al-Shehbaz: Harvard Pap. Bot. 12: 249. (2007) Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 263. (2007)
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