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Tulare rockcress

Short's rockcress

Habit Biennials or perennials; short-lived; apomictic; caudex present or absent. Biennials; short-lived; sexual; caudex present or absent.
Stems

usually 1 per caudex branch, arising from center of rosette near ground surface, 2–7 dm, sparsely to densely pubescent proximally, trichomes subsessile, sub-malpighiaceous, 0.3–0.6 mm, glabrous distally.

usually 2–4 per plant or caudex branch, arising from margin of rosette near ground surface, or arising laterally proximal to sterile shoots, 2–6 dm, pubescent proximally, trichomes short-stalked, 2–4-rayed, 0.15–0.4 mm, sparsely pubescent or glabrous distally.

Basal leaves

blade oblanceolate, 3–7 mm wide, margins entire, ciliate proximally, trichomes (simple), to 0.8 mm, surfaces sparsely to densely pubescent, trichomes subsessile, 2–5-rayed, 0.2–0.55 mm.

blade obovate to oblanceolate, (6–)10–45 mm wide, margins dentate, not ciliate, surfaces sparsely pubescent, abaxially trichomes short-stalked, 3- or 4-rayed, 0.15–0.4 mm, adaxially simple ones, 0.4–1 mm.

Cauline leaves

7–17, often concealing stem proximally;

blade auricles 2–5 mm, surfaces of distalmost leaves sparsely pubescent or glabrous.

4–20, rarely concealing stem;

blade auricles 1–8 mm, surfaces of distalmost leaves sparsely pubescent.

Racemes

19–39-flowered, usually unbranched.

14–66-flowered, sparingly branched.

Flowers

divaricate to pendent at anthesis;

sepals pubescent;

petals white to pale lavender, 6–7 × 1.2–2 mm, glabrous;

pollen spheroid.

divaricate at anthesis;

sepals glabrous or sparsely pubescent;

petals white or cream, 2–3.5 × 0.7–1 mm, glabrous;

pollen ellipsoid.

Fruiting pedicels

reflexed, recurved proximally, 5–13 mm, glabrous.

divaricate-ascending to slightly descending, straight, 1–6 mm, glabrous or pubescent, trichomes appressed, branched.

Fruits

reflexed, rarely appressed to rachis, not secund, straight, edges parallel, 4–7(–8.5) cm × 2–2.3 mm;

valves glabrous;

ovules 88–104 per ovary;

style 0.3–0.7 mm.

divaricate-ascending to slightly descending, not appressed to rachis, not secund, straight or slightly curved, edges parallel, 1.3–4.2 cm × 0.7–0.8 mm;

valves usually sparsely pubescent throughout, rarely glabrous;

ovules 14–44 per ovary;

style 0.1–0.5 mm.

Seeds

sub-biseriate, 2–2.5 × 1.1–1.5 mm;

wing continuous, 0.15–0.25 mm wide.

uniseriate, 0.7–1.1 × 0.5–0.6 mm;

not winged.

Boechera tularensis

Boechera dentata

Phenology Flowering Jun–Jul. Flowering Apr–Jun.
Habitat Rocky slopes in montane and subalpine habitats Bluffs and rocky ledges, wooded slopes, floodplains
Elevation 2400-3200 m (7900-10500 ft) 100-300 m (300-1000 ft)
Distribution
from FNA
CA
[BONAP county map]
from FNA
AL; AR; DC; IA; IL; IN; KS; KY; MD; MI; MN; MO; NE; NY; OH; OK; PA; SD; TN; VA; WI; WV; ON
[WildflowerSearch map]
Discussion

Morphological evidence suggests that Boechera tularensis is an apomictic species that contains three different genomes, one each from B. rectissima, B. retrofracta, and B. stricta. It is most often confused with B. pinetorum and B. retrofracta, but is easily distinguishable from both (see M. D. Windham and I. A. Al-Shehbaz 2007b for detailed comparison). It is known from the southern Sierra Nevada (Fresno and Tulare counties).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Sisymbrium dentatum Torrey is an illegitimate name that pertains to Iodanthus dentatus.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 410. FNA vol. 7, p. 373.
Parent taxa Brassicaceae > tribe Boechereae > Boechera Brassicaceae > tribe Boechereae > Boechera
Sibling taxa
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
Synonyms Shortia dentata, Arabis dentata var. phalacrocarpa, Arabis perstellata var. shortii, Arabis shortii, Arabis shortii var. phalacrocarpa, B. shortii, Iodanthus dentatus
Name authority Windham & Al-Shehbaz: Harvard Pap. Bot. 12: 249. (2007) (Rafinesque) Al-Shehbaz & Zarucchi: Harvard Pap. Bot. 13: 293. (2008)
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