Boechera subpinnatifida |
Boechera pallidifolia |
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ashy rock-cress, Klamath rockcress |
Gunnison County rockcress |
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Habit | Perennials; long-lived; sexual; caudex woody (often with persistent, crowded leaf bases). | Perennials; short-lived; sexual; caudex usually not woody. |
Stems | usually 1 per caudex branch, arising from center of rosette near ground surface, 1–4(–5) dm, densely pubescent proximally, trichomes short-stalked, 2–6-rayed, 0.1–0.2 mm, sparsely pubescent distally. |
usually 2–5 per caudex branch, arising from margin of rosette near ground surface, or arising laterally proximal to sterile shoots, (0.5–)1.5–4 dm, densely pubescent proximally, trichomes short-stalked, 2–6-rayed, 0.1–0.3 mm, glabrous or sparsely pubescent distally. |
Basal leaves | blade narrowly oblanceolate, 1–4(–5) mm wide, margins prominently dentate to subpinnatifid (leaf margins of sterile shoots often entire), ciliate near petiole base, trichomes (simple or 2-rayed), 0.4–0.6 mm, surfaces densely pubescent, trichomes short-stalked, (2–)4–9-rayed, 0.05–0.2 mm. |
blade oblanceolate to obovate, 5–11(–13) mm wide, margins shallowly dentate or, sometimes, entire, often ciliate along petiole, trichomes (simple and 2-rayed), surfaces moderately pubescent, trichomes short-stalked, 4–8-rayed, 0.1–0.3 mm. |
Cauline leaves | (10–)20–60, often concealing stem throughout; blade auricles 0.5–3 mm, surfaces of distalmost leaves moderately to sparsely pubescent. |
3–8(–11), not concealing stem; blade auricles (0.5–)1–2 mm, surfaces of distalmost leaves sparsely pubescent. |
Racemes | 8–30-flowered, usually unbranched. |
4–15(–20)-flowered, usually unbranched. |
Flowers | divaricate-ascending to pendent at anthesis; sepals pubescent; petals usually purple, rarely lavender, 9–14 × 1.5–3 mm, glabrous; pollen ellipsoid. |
ascending at anthesis; sepals pubescent; petals usually lavender, rarely whitish, 5–9 × 1.2–2.5 mm, glabrous; pollen ellipsoid. |
Fruiting pedicels | reflexed, strongly recurved, 5–15 mm, pubescent, trichomes appressed, branched. |
ascending to divaricate-ascending, straight or, sometimes, recurved, 7–15 mm, glabrous or sparsely pubescent. |
Fruits | pendent, not appressed to rachis, not secund, straight to slightly curved, edges parallel, (3.5–)5–8 cm × (1.6–)2–3 mm; valves pubescent throughout; ovules 24–42 per ovary; style 0.5–1 mm. |
usually ascending, rarely horizontal, not appressed to rachis, not secund, straight to curved, edges parallel, 2.5–5(–6.5) cm × 1–2 mm; valves glabrous; ovules 44–62 per ovary; style 0.05–0.4 mm. |
Seeds | uniseriate, 2.5–3.5 × 1.5–2.2 mm; wing continuous or at both ends, 0.4–0.8 mm wide. |
uniseriate, 1.1–1.3 × 1–1.1 mm; wing nearly continuous, 0.07–0.15 mm wide. |
2n | = 14. |
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Boechera subpinnatifida |
Boechera pallidifolia |
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Phenology | Flowering Mar–May. | Flowering Apr–Jun. |
Habitat | Rock outcrops, talus, gravelly soil, often in sagebrush-grassland communities | Rocky slopes and sandy soil in pinyon-juniper, sagebrush communities |
Elevation | 800-2400 m (2600-7900 ft) | 1600-2500 m (5200-8200 ft) |
Distribution |
CA; ID; NV; OR; UT
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AZ; CO; NM; UT; WY
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Discussion | Originally thought to be restricted to northern California and adjacent Oregon, Boechera subpinnatifida is a sexual species that recently has been found in central Idaho, northern Nevada, and northwestern Utah. It appears to intergrade with both B. puberula and B. retrofracta, and species boundaries within this complex need further study. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Although Boechera pallidifolia was originally described as endemic to west-central Colorado, recent studies favor a broader circumscription that includes sexual populations previously assigned to Arabis (Boechera) selbyi (see M. D. Windham and I. A. Al-Shehbaz 2006 for detailed comparison). The species has hybridized with nearly every sexual diploid within its range (including B. crandallii, B. fendleri, B. formosa, and B. pendulina), producing a confusing array of apomictic triploids including the type specimen of A. selbyi. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 409. | FNA vol. 7, p. 392. |
Parent taxa | Brassicaceae > tribe Boechereae > Boechera | Brassicaceae > tribe Boechereae > Boechera |
Sibling taxa | ||
Synonyms | Arabis subpinnatifida | Arabis pallidifolia, Arabis thompsonii, B. thompsonii |
Name authority | (S. Watson) Al-Shehbaz: Novon 13: 389. (2003) | (Rollins) W. A. Weber: Phytologia 79: 65. (1996) |
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