Boechera subpinnatifida |
Boechera suffrutescens |
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ashy rock-cress, Klamath rockcress |
woody rock-cress |
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Habit | Perennials; long-lived; sexual; caudex woody (often with persistent, crowded leaf bases). | Perennials; long-lived; sexual or apomictic; caudex woody. |
Stems | usually 1 per caudex branch, arising from center of rosette near ground surface, 1–4(–5) dm, densely pubescent proximally, trichomes short-stalked, 2–6-rayed, 0.1–0.2 mm, sparsely pubescent distally. |
usually 1 per caudex branch, arising from center of rosette, elevated above ground surface on woody base, 1–4(–5) dm, usually glabrous proximally, rarely pubescent, trichomes short-stalked, 2-rayed, 0.1–0.3 mm, glabrous distally. |
Basal leaves | blade narrowly oblanceolate, 1–4(–5) mm wide, margins prominently dentate to subpinnatifid (leaf margins of sterile shoots often entire), ciliate near petiole base, trichomes (simple or 2-rayed), 0.4–0.6 mm, surfaces densely pubescent, trichomes short-stalked, (2–)4–9-rayed, 0.05–0.2 mm. |
blade narrowly oblanceolate, 1.5–6 mm wide, margins entire, not ciliate or, rarely, with simple trichomes to 0.7 mm, surfaces glabrous or sparsely to densely pubescent, trichomes short-stalked, 2–6-rayed, 0.07–0.4 mm. |
Cauline leaves | (10–)20–60, often concealing stem throughout; blade auricles 0.5–3 mm, surfaces of distalmost leaves moderately to sparsely pubescent. |
(4–)6–12, sometimes concealing stem proximally; blade auricles 0.5–2 mm, surfaces of distalmost leaves glabrous. |
Racemes | 8–30-flowered, usually unbranched. |
6–12-flowered, usually unbranched. |
Flowers | divaricate-ascending to pendent at anthesis; sepals pubescent; petals usually purple, rarely lavender, 9–14 × 1.5–3 mm, glabrous; pollen ellipsoid. |
pendent at anthesis; sepals glabrous; petals purple or whitish with rose tips, 4.5–6 × 2–2.5 mm, glabrous; pollen ellipsoid or spheroid. |
Fruiting pedicels | reflexed, strongly recurved, 5–15 mm, pubescent, trichomes appressed, branched. |
reflexed, usually abruptly recurved at base, otherwise straight, 4–18 mm, glabrous. |
Fruits | pendent, not appressed to rachis, not secund, straight to slightly curved, edges parallel, (3.5–)5–8 cm × (1.6–)2–3 mm; valves pubescent throughout; ovules 24–42 per ovary; style 0.5–1 mm. |
reflexed to closely pendent, sometimes appressed to rachis, often secund, straight to somewhat curved, edges undulate (not parallel), 3–7(–8) cm × 3–5(–6) mm; valves glabrous; ovules 20–30 per ovary; style 0.4–1.2 mm. |
Seeds | uniseriate, 2.5–3.5 × 1.5–2.2 mm; wing continuous or at both ends, 0.4–0.8 mm wide. |
uniseriate, 2.5–5.5 × 1.8–3.5 mm; wing continuous, 0.8–1.5 mm wide. |
2n | = 14. |
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Boechera subpinnatifida |
Boechera suffrutescens |
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Phenology | Flowering Mar–May. | Flowering Jun–Aug. |
Habitat | Rock outcrops, talus, gravelly soil, often in sagebrush-grassland communities | Rocky slopes and gravelly soil, often with sagebrush |
Elevation | 800-2400 m (2600-7900 ft) | 1800-3000 m (5900-9800 ft) |
Distribution |
CA; ID; NV; OR; UT
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CA; ID; NV; OR; WA
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Discussion | Originally thought to be restricted to northern California and adjacent Oregon, Boechera subpinnatifida is a sexual species that recently has been found in central Idaho, northern Nevada, and northwestern Utah. It appears to intergrade with both B. puberula and B. retrofracta, and species boundaries within this complex need further study. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Boechera suffrutescens is recognizable by its prominently suffrutescent habit and wide (greater than 3 mm), reflexed fruits. Both sexual and apomictic collections are known; further study is needed to determine whether they truly are conspecific. The taxon previously known as Arabis suffrutescens var. horizontalis appears to be of hybrid origin; it is treated here as a distinct species (see M. D. Windham and I. A. Al-Shehbaz 2007 for detailed comparison). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 409. | FNA vol. 7, p. 409. |
Parent taxa | Brassicaceae > tribe Boechereae > Boechera | Brassicaceae > tribe Boechereae > Boechera |
Sibling taxa | ||
Synonyms | Arabis subpinnatifida | Arabis suffrutescens, Arabis dianthifolia, Arabis duriuscula |
Name authority | (S. Watson) Al-Shehbaz: Novon 13: 389. (2003) | (S. Watson) Dorn: Brittonia 55: 3. (2003) |
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