Boechera subpinnatifida |
Boechera platysperma |
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ashy rock-cress, Klamath rockcress |
broad-seed rock-cress, flat-seed rock-cress, pioneer rock-cress |
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Habit | Perennials; long-lived; sexual; caudex woody (often with persistent, crowded leaf bases). | Perennials; long-lived; sexual; caudex woody. |
Stems | usually 1 per caudex branch, arising from center of rosette near ground surface, 1–4(–5) dm, densely pubescent proximally, trichomes short-stalked, 2–6-rayed, 0.1–0.2 mm, sparsely pubescent distally. |
usually 1 per caudex branch, arising from center of rosette, elevated on woody base above or near ground surface, 0.6–2.8(–3.5) dm, densely pubescent proximally, trichomes short-stalked, 2–5-rayed, 0.1–0.3 mm, glabrous or sparsely pubescent distally. |
Basal leaves | blade narrowly oblanceolate, 1–4(–5) mm wide, margins prominently dentate to subpinnatifid (leaf margins of sterile shoots often entire), ciliate near petiole base, trichomes (simple or 2-rayed), 0.4–0.6 mm, surfaces densely pubescent, trichomes short-stalked, (2–)4–9-rayed, 0.05–0.2 mm. |
blade narrowly oblanceolate, 3–7(–10) mm wide, margins entire, ciliate near petiole base, trichomes (simple or spurred), to 1 mm, surfaces densely pubescent, trichomes short-stalked, 2–4 (or 5)-rayed 0.1–0.3 mm. |
Cauline leaves | (10–)20–60, often concealing stem throughout; blade auricles 0.5–3 mm, surfaces of distalmost leaves moderately to sparsely pubescent. |
3–7(–12), not concealing stem; blade auricles absent, surfaces of distalmost leaves glabrous or sparsely pubescent. |
Racemes | 8–30-flowered, usually unbranched. |
2–7-flowered, unbranched. |
Flowers | divaricate-ascending to pendent at anthesis; sepals pubescent; petals usually purple, rarely lavender, 9–14 × 1.5–3 mm, glabrous; pollen ellipsoid. |
ascending at anthesis; sepals pubescent; petals creamy white to purplish, 4–6 × 1–2 mm, sparsely pubescent (often some trichomes abaxially); pollen ellipsoid. |
Fruiting pedicels | reflexed, strongly recurved, 5–15 mm, pubescent, trichomes appressed, branched. |
ascending, straight, 3–13 mm, glabrous or sparsely pubescent, trichomes appressed, branched. |
Fruits | pendent, not appressed to rachis, not secund, straight to slightly curved, edges parallel, (3.5–)5–8 cm × (1.6–)2–3 mm; valves pubescent throughout; ovules 24–42 per ovary; style 0.5–1 mm. |
ascending, not appressed to rachis, not secund, straight to slightly curved, edges undulate (not parallel), (2.5–)4–8.5 cm × 3–5.5 mm; valves glabrous; ovules 16–44 per ovary; style 0.05–1 mm. |
Seeds | uniseriate, 2.5–3.5 × 1.5–2.2 mm; wing continuous or at both ends, 0.4–0.8 mm wide. |
uniseriate, 3–6(–8) × 2–3.5 mm; wing continuous, 0.8–2.5 mm wide. |
2n | = 14. |
= 14. |
Boechera subpinnatifida |
Boechera platysperma |
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Phenology | Flowering Mar–May. | Flowering Jun–Aug. |
Habitat | Rock outcrops, talus, gravelly soil, often in sagebrush-grassland communities | Rock outcrops and gravelly soil in dry pine forests and lodgepole-chaparral woodlands |
Elevation | 800-2400 m (2600-7900 ft) | 1600-3000 m (5200-9800 ft) |
Distribution |
CA; ID; NV; OR; UT
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CA; OR
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Discussion | Originally thought to be restricted to northern California and adjacent Oregon, Boechera subpinnatifida is a sexual species that recently has been found in central Idaho, northern Nevada, and northwestern Utah. It appears to intergrade with both B. puberula and B. retrofracta, and species boundaries within this complex need further study. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Previous authors have adopted broader circumscriptions of Boechera platysperma, including plants herein called B. covillei, B. elkoensis, and B. howellii. In our opinion, the differences between these taxa are sufficient to justify their treatment as distinct species (see M. D. Windham and I. A. Al-Shehbaz 2006, 2007, 2007b for detailed comparison). Reports of B. platysperma from Nevada (e.g., R. C. Rollins 1993; N. H. Holmgren 2005b) are based on plants of B. howellii or apomictic hybrids with other species. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 409. | FNA vol. 7, p. 397. |
Parent taxa | Brassicaceae > tribe Boechereae > Boechera | Brassicaceae > tribe Boechereae > Boechera |
Sibling taxa | ||
Synonyms | Arabis subpinnatifida | Arabis platysperma, Arabis inamoena, Arabis oligantha |
Name authority | (S. Watson) Al-Shehbaz: Novon 13: 389. (2003) | (A. Gray) Al-Shehbaz: Novon 13: 388. (2003) |
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