Boechera subpinnatifida |
Boechera pauciflora |
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ashy rock-cress, Klamath rockcress |
Columbia rockcress, few-flower rock-cress, fuzzy suncress, hairy-stem rock-cress, small-flower rockcress |
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Habit | Perennials; long-lived; sexual; caudex woody (often with persistent, crowded leaf bases). | Perennials; short- to long-lived; apomictic; caudex sometimes woody. |
Stems | usually 1 per caudex branch, arising from center of rosette near ground surface, 1–4(–5) dm, densely pubescent proximally, trichomes short-stalked, 2–6-rayed, 0.1–0.2 mm, sparsely pubescent distally. |
usually 1 per caudex branch, arising from center of rosette near ground surface, (1.4–)3–11.2 dm, densely pubescent proximally, trichomes simple, 0.6–1.5 mm, mixed with stalked, 2- (or 3-)rayed ones, 0.2–0.4 mm, sparsely pubescent distally. |
Basal leaves | blade narrowly oblanceolate, 1–4(–5) mm wide, margins prominently dentate to subpinnatifid (leaf margins of sterile shoots often entire), ciliate near petiole base, trichomes (simple or 2-rayed), 0.4–0.6 mm, surfaces densely pubescent, trichomes short-stalked, (2–)4–9-rayed, 0.05–0.2 mm. |
blade oblanceolate, 3–10 mm wide, margins usually dentate, rarely entire, sometimes ciliate, trichomes (simple or branched), to 1 mm, surfaces densely pubescent, trichomes stalked, 2–5-rayed, 0.3–0.6 mm. |
Cauline leaves | (10–)20–60, often concealing stem throughout; blade auricles 0.5–3 mm, surfaces of distalmost leaves moderately to sparsely pubescent. |
(8–)14–60, often concealing stem proximally; blade auricles (1–)3–10 mm, surfaces of distalmost leaves glabrous or sparsely pubescent. |
Racemes | 8–30-flowered, usually unbranched. |
17–60-flowered, usually unbranched. |
Flowers | divaricate-ascending to pendent at anthesis; sepals pubescent; petals usually purple, rarely lavender, 9–14 × 1.5–3 mm, glabrous; pollen ellipsoid. |
divaricate at anthesis; sepals pubescent; petals lavender to whitish, 5–8 × 1–2 mm, glabrous; pollen spheroid. |
Fruiting pedicels | reflexed, strongly recurved, 5–15 mm, pubescent, trichomes appressed, branched. |
horizontal to divaricate-descending, usually straight, rarely slightly recurved, 4–13 mm, pubescent, trichomes spreading, 2- or 3-rayed. |
Fruits | pendent, not appressed to rachis, not secund, straight to slightly curved, edges parallel, (3.5–)5–8 cm × (1.6–)2–3 mm; valves pubescent throughout; ovules 24–42 per ovary; style 0.5–1 mm. |
horizontal, divaricate-descending or widely pendent, not appressed to rachis, not secund, curved, edges parallel, 5.5–10.5 cm × 1.5–2.2 mm; valves glabrous; ovules 80–162 per ovary; style 0.05–0.5 mm. |
Seeds | uniseriate, 2.5–3.5 × 1.5–2.2 mm; wing continuous or at both ends, 0.4–0.8 mm wide. |
uniseriate, 1.4–1.8 × 1–1.4 mm; wing continuous, 0.1–0.25 mm wide. |
2n | = 14. |
= 21. |
Boechera subpinnatifida |
Boechera pauciflora |
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Phenology | Flowering Mar–May. | Flowering May–Jun. |
Habitat | Rock outcrops, talus, gravelly soil, often in sagebrush-grassland communities | Rocky soil in sagebrush areas, mountain shrub communities, edges of conifer forests |
Elevation | 800-2400 m (2600-7900 ft) | 600-2500 m (2000-8200 ft) |
Distribution |
CA; ID; NV; OR; UT
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CA; ID; MT; NV; OR; UT; WA; WY; BC
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Discussion | Originally thought to be restricted to northern California and adjacent Oregon, Boechera subpinnatifida is a sexual species that recently has been found in central Idaho, northern Nevada, and northwestern Utah. It appears to intergrade with both B. puberula and B. retrofracta, and species boundaries within this complex need further study. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Morphological evidence suggests that Boechera pauciflora is an apomictic species that arose through hybridization between B. retrofracta and B. sparsiflora. Specimens of B. pauciflora are commonly identified as Arabis holboellii var. pinetorum (= B. pinetorum), a superficially similar species restricted to the northern Sierra Nevada and southern Cascade Range (see M. D. Windham and I. A. 2007 for detailed comparison). Arabis elegans A. Nelson (1900), not Tineo & Lojacono (1886) is an illegitimate name, sometimes found in synonymy with Boechera pauciflora. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 409. | FNA vol. 7, p. 393. |
Parent taxa | Brassicaceae > tribe Boechereae > Boechera | Brassicaceae > tribe Boechereae > Boechera |
Sibling taxa | ||
Synonyms | Arabis subpinnatifida | Sisymbrium pauciflorum, Arabis arcuata var. subvillosa, Arabis columbiana, Arabis perelegans, Arabis sparsiflora var. columbiana, Arabis sparsiflora var. subvillosa, B. sparsiflora var. subvillosa |
Name authority | (S. Watson) Al-Shehbaz: Novon 13: 389. (2003) | (Nuttall) Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 268. (2007) |
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