Boechera subpinnatifida |
Boechera microphylla |
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ashy rock-cress, Klamath rockcress |
crevice suncress, little-leaf rockcress, small-flower rockcress, small-leaf rock-cress |
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Habit | Perennials; long-lived; sexual; caudex woody (often with persistent, crowded leaf bases). | Perennials; long-lived; (cespitose); sexual or apomictic; caudex woody. |
Stems | usually 1 per caudex branch, arising from center of rosette near ground surface, 1–4(–5) dm, densely pubescent proximally, trichomes short-stalked, 2–6-rayed, 0.1–0.2 mm, sparsely pubescent distally. |
1 or several per caudex branch, arising from center or margin of rosette near ground surface, 0.8–2.8(–3.5) dm, sparsely pubescent proximally, trichomes simple and subsessile, 2-rayed, 0.3–0.6 mm, glabrous distally. |
Basal leaves | blade narrowly oblanceolate, 1–4(–5) mm wide, margins prominently dentate to subpinnatifid (leaf margins of sterile shoots often entire), ciliate near petiole base, trichomes (simple or 2-rayed), 0.4–0.6 mm, surfaces densely pubescent, trichomes short-stalked, (2–)4–9-rayed, 0.05–0.2 mm. |
blade oblanceolate to linear-oblanceolate, 1–4(–6) mm wide, margins entire or subapically toothed, ciliate proximally, trichomes (simple), to 0.6 mm, surfaces densely pubescent, trichomes short-stalked, 4–8-rayed, 0.05–0.1 mm. |
Cauline leaves | (10–)20–60, often concealing stem throughout; blade auricles 0.5–3 mm, surfaces of distalmost leaves moderately to sparsely pubescent. |
2–6, not concealing stem; blade auricles 0.5–1.5 mm, surfaces of distalmost leaves glabrous. |
Racemes | 8–30-flowered, usually unbranched. |
5–17-flowered, usually unbranched. |
Flowers | divaricate-ascending to pendent at anthesis; sepals pubescent; petals usually purple, rarely lavender, 9–14 × 1.5–3 mm, glabrous; pollen ellipsoid. |
ascending at anthesis; sepals glabrous or pubescent; petals lavender to purple, 3.5–5.5 × 1–1.8 mm, glabrous; pollen ellipsoid or spheroid. |
Fruiting pedicels | reflexed, strongly recurved, 5–15 mm, pubescent, trichomes appressed, branched. |
ascending to divaricate-ascending, straight, 4–14 mm, glabrous. |
Fruits | pendent, not appressed to rachis, not secund, straight to slightly curved, edges parallel, (3.5–)5–8 cm × (1.6–)2–3 mm; valves pubescent throughout; ovules 24–42 per ovary; style 0.5–1 mm. |
ascending to divaricate-ascending, not appressed to rachis, not secund, straight to slightly curved, edges parallel, 3–7 cm × 1–1.5 mm; valves glabrous; ovules 52–72 per ovary; style 0.05–0.3 mm. |
Seeds | uniseriate, 2.5–3.5 × 1.5–2.2 mm; wing continuous or at both ends, 0.4–0.8 mm wide. |
uniseriate, 1–1.5 × 0.7–1 mm; wing often distal, ca. 0.1 mm wide. |
2n | = 14. |
= 14, 15. |
Boechera subpinnatifida |
Boechera microphylla |
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Phenology | Flowering Mar–May. | Flowering Apr–Jun. |
Habitat | Rock outcrops, talus, gravelly soil, often in sagebrush-grassland communities | Cliffs and rocky slopes in sagebrush, mountain shrub, and open conifer forests |
Elevation | 800-2400 m (2600-7900 ft) | 400-2400 m (1300-7900 ft) |
Distribution |
CA; ID; NV; OR; UT
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ID; MT; NV; OR; UT; WA; WY; BC
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Discussion | Originally thought to be restricted to northern California and adjacent Oregon, Boechera subpinnatifida is a sexual species that recently has been found in central Idaho, northern Nevada, and northwestern Utah. It appears to intergrade with both B. puberula and B. retrofracta, and species boundaries within this complex need further study. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Boechera microphylla is recognizable by its minute (0.05–0.1 mm), 4–8-rayed leaf trichomes, mat-forming habit, simple and 2-rayed trichomes on stems proximally, and ascending fruits. Both sexual and apomictic collections are known; further study is needed to determine whether they truly are conspecific. The taxa traditionally treated as Arabis (Boechera) microphylla vars. macounii and thompsonii are here recognized as B. macounii and B. cascadensis, respectively (see M. D. Windham and I. A. Al-Shehbaz 2007 for detailed comparison). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 409. | FNA vol. 7, p. 390. |
Parent taxa | Brassicaceae > tribe Boechereae > Boechera | Brassicaceae > tribe Boechereae > Boechera |
Sibling taxa | ||
Synonyms | Arabis subpinnatifida | Arabis microphylla, Arabis tenuicula |
Name authority | (S. Watson) Al-Shehbaz: Novon 13: 389. (2003) | (Nuttall) Dorn: Vasc. Pl. Wyoming ed. 3, 376. (2001) |
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