Boechera subpinnatifida |
Boechera cobrensis |
|
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ashy rock-cress, Klamath rockcress |
cobre rock-cress, Masonic rock-cress, sagebrush rockcress |
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Habit | Perennials; long-lived; sexual; caudex woody (often with persistent, crowded leaf bases). | Perennials; long-lived; (often cespitose); sexual; caudex somewhat woody (usually without persistent leaf bases). |
Stems | usually 1 per caudex branch, arising from center of rosette near ground surface, 1–4(–5) dm, densely pubescent proximally, trichomes short-stalked, 2–6-rayed, 0.1–0.2 mm, sparsely pubescent distally. |
usually 1 per caudex branch, arising from center of rosette near ground surface, (1.2–)2.5–6 dm, densely pubescent proximally, trichomes short-stalked, 4–10-rayed, 0.1–0.2 mm, sparsely pubescent or glabrous distally. |
Basal leaves | blade narrowly oblanceolate, 1–4(–5) mm wide, margins prominently dentate to subpinnatifid (leaf margins of sterile shoots often entire), ciliate near petiole base, trichomes (simple or 2-rayed), 0.4–0.6 mm, surfaces densely pubescent, trichomes short-stalked, (2–)4–9-rayed, 0.05–0.2 mm. |
blade linear to narrowly oblanceolate, 1–4 mm wide, margins entire, ciliate near petiole base, trichomes (simple), to 0.6 mm, surfaces densely pubescent, trichomes short-stalked, 4–10-rayed, 0.1–0.2 mm. |
Cauline leaves | (10–)20–60, often concealing stem throughout; blade auricles 0.5–3 mm, surfaces of distalmost leaves moderately to sparsely pubescent. |
5–10, not concealing stem; blade auricles 1–1.5 mm, surfaces of distalmost leaves usually sparsely pubescent. |
Racemes | 8–30-flowered, usually unbranched. |
10–25-flowered, often sparingly branched. |
Flowers | divaricate-ascending to pendent at anthesis; sepals pubescent; petals usually purple, rarely lavender, 9–14 × 1.5–3 mm, glabrous; pollen ellipsoid. |
divaricate-ascending at anthesis; sepals pubescent; petals white to lavender, 3.5–6 × 0.7–1 mm, glabrous; pollen ellipsoid. |
Fruiting pedicels | reflexed, strongly recurved, 5–15 mm, pubescent, trichomes appressed, branched. |
divaricate-ascending to horizontal, often straight proximally, usually strongly recurved or reflexed distally, 4–17 mm, usually pubescent, rarely glabrous, trichomes appressed, branched. |
Fruits | pendent, not appressed to rachis, not secund, straight to slightly curved, edges parallel, (3.5–)5–8 cm × (1.6–)2–3 mm; valves pubescent throughout; ovules 24–42 per ovary; style 0.5–1 mm. |
pendent, not appressed to rachis, not secund, straight or slightly curved, edges parallel, 2.5–5.5 cm × 1.7–2.5 mm; valves glabrous; ovules 34–64 per ovary; style 0.05–0.2 mm. |
Seeds | uniseriate, 2.5–3.5 × 1.5–2.2 mm; wing continuous or at both ends, 0.4–0.8 mm wide. |
uniseriate, 1.4–1.8 × 1–1.2 mm; wing continuous, 0.25–0.5 mm wide. |
2n | = 14. |
= 14. |
Boechera subpinnatifida |
Boechera cobrensis |
|
Phenology | Flowering Mar–May. | Flowering May–Jun. |
Habitat | Rock outcrops, talus, gravelly soil, often in sagebrush-grassland communities | Sandy soil, usually under shelter of shrubs in semi-desert communities |
Elevation | 800-2400 m (2600-7900 ft) | 1200-2800 m (3900-9200 ft) |
Distribution |
CA; ID; NV; OR; UT
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CA; ID; NV; OR; WY
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Discussion | Originally thought to be restricted to northern California and adjacent Oregon, Boechera subpinnatifida is a sexual species that recently has been found in central Idaho, northern Nevada, and northwestern Utah. It appears to intergrade with both B. puberula and B. retrofracta, and species boundaries within this complex need further study. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Boechera cobrensis is a sexual diploid commonly found on stabilized sand dunes. The related B. lignifera often is found in similar habitats, and the two species can be difficult to distinguish where their ranges overlap in Idaho and Wyoming. It is possible that B. cobrensis was involved in the origin of some apomictic populations currently assigned to B. lignifera. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 409. | FNA vol. 7, p. 370. |
Parent taxa | Brassicaceae > tribe Boechereae > Boechera | Brassicaceae > tribe Boechereae > Boechera |
Sibling taxa | ||
Synonyms | Arabis subpinnatifida | Arabis cobrensis |
Name authority | (S. Watson) Al-Shehbaz: Novon 13: 389. (2003) | (M. E. Jones) Dorn: Vasc. Pl. Wyoming ed. 3, 375. (2001) |
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