Boechera spatifolia |
Boechera hastatula |
|
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spoonleaf rockcress |
Hells Canyon rock-cress |
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Habit | Perennials; short- to long-lived; sexual; caudex usually not woody (rarely with persistent, crowded leaf bases). | Perennials; short- to long-lived; apomictic or sexual; caudex often woody. |
Stems | simple or few to several per caudex branch, arising from center of rosette near ground surface, 1.5–3.5(–5) dm, densely pubescent proximally, trichomes simple mixed with short-stalked ones, 2-rayed, 0.3–0.7 mm. |
1–4 per caudex branch, arising from margin of rosette near ground surface, 1–3 dm, glabrous throughout. |
Basal leaves | blade narrowly oblanceolate, 1.5–3(–4) mm wide, margins entire, strongly ciliate at least along petiole, trichomes (simple), to 1 mm, surfaces glabrous or sparsely pubescent, trichomes simple and short-stalked, 2-rayed, 0.3–0.7 mm. |
blade linear-oblanceolate to oblanceolate, 1–3(–5) mm wide, margins entire or minutely toothed, rarely ciliate near petiole base, trichomes (simple), to 0.8 mm, surfaces pubescent or glabrous, trichomes short-stalked, 3–6-rayed, 0.07–0.2 mm. |
Cauline leaves | 5–15(–20), often concealing stem proximally; blade auricles 0.5–1.5 mm, surfaces of distalmost leaves usually glabrous. |
4–15, not concealing stem; blade auricles 0.5–2.5 mm, surfaces of distalmost leaves glabrous. |
Racemes | 10–30-flowered, usually unbranched. |
12–40-flowered, usually unbranched. |
Flowers | divaricate-ascending at anthesis; sepals pubescent; petals usually white, rarely pale lavender, 3–3.7(–4) × 0.5–0.8 mm, glabrous; pollen ellipsoid. |
divaricate to pendent at anthesis; sepals pubescent; petals lavender, 5–6 × ca. 1 mm, glabrous; pollen ellipsoid or spheroid. |
Fruiting pedicels | horizontal or slightly descending, curved or angled downward, 6–10(–15) mm, usually glabrous, rarely sparsely pubescent, trichomes spreading, simple. |
reflexed, abruptly recurved near base, 4–8 mm, glabrous. |
Fruits | pendent, not appressed to rachis, rarely slightly secund, straight or gently curved, edges parallel, 3.3–5.7 cm × 1.2–1.8 mm; valves glabrous; ovules 90–126 per ovary; style 0.1–0.4 mm. |
reflexed to closely pendent, usually appressed to rachis, often secund, straight, edges parallel, 0.3–0.42 cm × 1–1.5 mm; valves glabrous; ovules 46–60 per ovary. |
Seeds | biseriate, 0.7–0.9 × 0.5–0.6 mm; wing distal or, sometimes, absent, 0.05–0.1 mm wide. |
uniseriate, 1–1.3 × 0.8–1 mm; wing continuous, 0.05–0.1 mm wide. |
2n | = 14. |
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Boechera spatifolia |
Boechera hastatula |
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Phenology | Flowering May–Aug. | Flowering Jun–Jul. |
Habitat | Rocky slopes and gravelly soil in sagebrush, pinyon-juniper woodlands, open conifer forests and subalpine meadows | Igneous rock outcrops |
Elevation | 1800-2800 m (5900-9200 ft) | 1900-2100 m (6200-6900 ft) |
Distribution |
CO; NM
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OR |
Discussion | Boechera spatifolia is a sexual diploid that usually has been treated as a variety of Arabis (Boechera) fendleri but appears to be sufficiently distinct to warrant recognition at species level (see M. D. Windham and I. A. Al-Shehbaz 2006 for detailed comparison). There is little geographic overlap between the two, with B. spatifolia confined to the mountains of central Colorado and north-central New Mexico and B. fendleri ranging from western New Mexico and the Four Corners region through northern Arizona to southern Nevada. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Both sexual and apomictic collections of Boechera hastatula have been identified; further study is needed to determine whether they truly are conspecific. The species is known only from Baker and Wallowa counties. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 408. | FNA vol. 7, p. 383. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Arabis spatifolia, Arabis fendleri var. spatifolia, B. fendleri subsp. spatifolia, B. fendleri var. spatifolia | Arabis hastatula |
Name authority | (Rydberg) Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 84. (2006) | (Greene) Al-Shehbaz: Novon 13: 386. (2003) |
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