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spoonleaf rockcress

Hells Canyon rock-cress

Habit Perennials; short- to long-lived; sexual; caudex usually not woody (rarely with persistent, crowded leaf bases). Perennials; short- to long-lived; apomictic or sexual; caudex often woody.
Stems

simple or few to several per caudex branch, arising from center of rosette near ground surface, 1.5–3.5(–5) dm, densely pubescent proximally, trichomes simple mixed with short-stalked ones, 2-rayed, 0.3–0.7 mm.

1–4 per caudex branch, arising from margin of rosette near ground surface, 1–3 dm, glabrous throughout.

Basal leaves

blade narrowly oblanceolate, 1.5–3(–4) mm wide, margins entire, strongly ciliate at least along petiole, trichomes (simple), to 1 mm, surfaces glabrous or sparsely pubescent, trichomes simple and short-stalked, 2-rayed, 0.3–0.7 mm.

blade linear-oblanceolate to oblanceolate, 1–3(–5) mm wide, margins entire or minutely toothed, rarely ciliate near petiole base, trichomes (simple), to 0.8 mm, surfaces pubescent or glabrous, trichomes short-stalked, 3–6-rayed, 0.07–0.2 mm.

Cauline leaves

5–15(–20), often concealing stem proximally;

blade auricles 0.5–1.5 mm, surfaces of distalmost leaves usually glabrous.

4–15, not concealing stem;

blade auricles 0.5–2.5 mm, surfaces of distalmost leaves glabrous.

Racemes

10–30-flowered, usually unbranched.

12–40-flowered, usually unbranched.

Flowers

divaricate-ascending at anthesis;

sepals pubescent;

petals usually white, rarely pale lavender, 3–3.7(–4) × 0.5–0.8 mm, glabrous;

pollen ellipsoid.

divaricate to pendent at anthesis;

sepals pubescent;

petals lavender, 5–6 × ca. 1 mm, glabrous;

pollen ellipsoid or spheroid.

Fruiting pedicels

horizontal or slightly descending, curved or angled downward, 6–10(–15) mm, usually glabrous, rarely sparsely pubescent, trichomes spreading, simple.

reflexed, abruptly recurved near base, 4–8 mm, glabrous.

Fruits

pendent, not appressed to rachis, rarely slightly secund, straight or gently curved, edges parallel, 3.3–5.7 cm × 1.2–1.8 mm;

valves glabrous;

ovules 90–126 per ovary;

style 0.1–0.4 mm.

reflexed to closely pendent, usually appressed to rachis, often secund, straight, edges parallel, 0.3–0.42 cm × 1–1.5 mm;

valves glabrous;

ovules 46–60 per ovary.

Seeds

biseriate, 0.7–0.9 × 0.5–0.6 mm;

wing distal or, sometimes, absent, 0.05–0.1 mm wide.

uniseriate, 1–1.3 × 0.8–1 mm;

wing continuous, 0.05–0.1 mm wide.

2n

= 14.

Boechera spatifolia

Boechera hastatula

Phenology Flowering May–Aug. Flowering Jun–Jul.
Habitat Rocky slopes and gravelly soil in sagebrush, pinyon-juniper woodlands, open conifer forests and subalpine meadows Igneous rock outcrops
Elevation 1800-2800 m (5900-9200 ft) 1900-2100 m (6200-6900 ft)
Distribution
from FNA
CO; NM
[WildflowerSearch map]
[BONAP county map]
from FNA
OR
[BONAP county map]
Discussion

Boechera spatifolia is a sexual diploid that usually has been treated as a variety of Arabis (Boechera) fendleri but appears to be sufficiently distinct to warrant recognition at species level (see M. D. Windham and I. A. Al-Shehbaz 2006 for detailed comparison). There is little geographic overlap between the two, with B. spatifolia confined to the mountains of central Colorado and north-central New Mexico and B. fendleri ranging from western New Mexico and the Four Corners region through northern Arizona to southern Nevada.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Both sexual and apomictic collections of Boechera hastatula have been identified; further study is needed to determine whether they truly are conspecific. The species is known only from Baker and Wallowa counties.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 408. FNA vol. 7, p. 383.
Parent taxa Brassicaceae > tribe Boechereae > Boechera Brassicaceae > tribe Boechereae > Boechera
Sibling taxa
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hoffmannii, B. holboellii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
Synonyms Arabis spatifolia, Arabis fendleri var. spatifolia, B. fendleri subsp. spatifolia, B. fendleri var. spatifolia Arabis hastatula
Name authority (Rydberg) Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 84. (2006) (Greene) Al-Shehbaz: Novon 13: 386. (2003)
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