Boechera sparsiflora |
Boechera horizontalis |
|
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elegant rockcress, sickle-pod rock-cress, slender rockcress, stretching suncress |
Crater Lake rock-cress |
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Habit | Biennials or perennials; short-lived; sexual; caudex present or absent. | Perennials; long-lived; (cespitose); apomictic; caudex woody. |
Stems | usually 1 per caudex branch, arising from center of rosette near ground surface, 3–8 dm, densely pubescent proximally, trichomes simple mixed with fewer short-stalked, 2-rayed ones, 0.4–1.5 mm, glabrous or sparsely pubescent distally. |
usually 1 per caudex branch, arising from center of rosette, near ground surface or somewhat elevated on woody base, or 1–3.5 dm, sparsely pubescent proximally, trichomes short-stalked, 3–6-rayed, 0.1–0.3 mm, glabrous distally. |
Basal leaves | blade oblanceolate or linear-oblanceolate, 3–12 mm wide, margins usually entire, rarely dentate, ciliate proximally, trichomes (usually simple), to 1 mm, surfaces densely pubescent, trichomes short-stalked, 2–5-rayed, 0.3–0.8 mm. |
blade narrowly oblanceolate, 1–5 mm wide, margins entire, ciliate near petiole base, trichomes (simple), to 0.4 mm, surfaces densely pubescent, trichomes short-stalked, 3–6-rayed, 0.1–0.3 mm. |
Cauline leaves | (9–)15–35, often concealing stem proximally; blade auricles 3–10 mm, surfaces of distalmost leaves glabrous, sometimes margins ciliate. |
3–13, often concealing stem proximally; blade auricles 0.5–1.5 mm, surfaces of distalmost leaves glabrous. |
Racemes | 12–50-flowered, usually unbranched. |
5–32-flowered, usually unbranched. |
Flowers | ascending at anthesis; sepals sparsely pubescent; petals usually lavender to purple, rarely white, 7–13 × 2–5 mm, glabrous; pollen ellipsoid. |
divaricate-ascending at anthesis; sepals pubescent; petals lavender to purple, 5–6 × 1.5–2 mm, glabrous; pollen spheroid. |
Fruiting pedicels | usually ascending, rarely almost horizontal, straight or slightly recurved, 3–10(–18) mm, usually pubescent, rarely glabrous, trichomes spreading, usually simple. |
horizontal to descending, straight or slightly curved downward, 4–11 mm, glabrous. |
Fruits | usually ascending, rarely almost horizontal, not appressed to rachis, not secund, usually curved, edges parallel, 5–13 cm × 1.7–2 mm; valves glabrous; ovules 90–170 per ovary; style 0.05–0.3 mm. |
horizontal or descending, not appressed to rachis, secund, straight, edges slightly undulate (not parallel), 2–4 cm × 2–3 mm; valves glabrous; ovules 40–54 per ovary; style 0.2–0.5 mm. |
Seeds | uniseriate, 1.5–2 × 1–1.5 mm; wing continuous, 0.1–0.2 mm wide. |
uniseriate, 2–2.5 × 1.7–2 mm; wing continuous, 0.5–1 mm wide. |
2n | = 14. |
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Boechera sparsiflora |
Boechera horizontalis |
|
Phenology | Flowering Apr–Jun. | Flowering Jul–Aug. |
Habitat | Rocky slopes, clay hills, sandy soil in sagebrush and mountain shrub communities, meadows, and open conifer forests | Dry pumice slopes |
Elevation | 400-2800 m (1300-9200 ft) | |
Distribution |
CA; ID; NV; OR; UT; WA; BC
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OR |
Discussion | As circumscribed by R. C. Rollins (1993), Boechera sparsiflora included six varieties encompassing three sexual diploids and a number of apomictic hybrids. The most distinctive of those elements are recognized here as the separate species B. arcuata, B. atrorubens, B. californica, and B. pauciflora. The narrow circumscription of B. sparsiflora adopted here includes only sexual diploids. It is distinguished from other taxa previously assigned to it by having proximal stems densely pubescent with predominantly simple (some 2-rayed) trichomes to 1.5 mm, usually glabrous distal stems, and ascending fruiting pedicels with spreading, usually simple trichomes (rarely glabrous). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Morphological evidence suggests that Boechera horizontalis is an apomictic species that arose through hybridization between B. lemmonii and B. suffrutescens (M. D. Windham and I. A. Al-Shehbaz 2007). Boechera horizontalis is known only from the vicinity of Crater Lake in south-central Oregon. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 407. | FNA vol. 7, p. 384. |
Parent taxa | Brassicaceae > tribe Boechereae > Boechera | Brassicaceae > tribe Boechereae > Boechera |
Sibling taxa | ||
Synonyms | Arabis sparsiflora, Arabis arcoidea, Arabis campyloloba, Arabis peramoena, Arabis polytricha, Arabis sparsiflora var. peramoena | Arabis horizontalis, Arabis suffrutescens var. horizontalis |
Name authority | (Nuttall) Dorn: Vasc. Pl. Wyoming ed. 3, 376. (2001) | (Greene) Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 266. (2007) |
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