Boechera shockleyi |
Boechera williamsii |
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Shockley's Rock cress |
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Habit | Perennials; usually short-lived; sexual; caudex not woody, (rarely with persistent, crowded leaf bases). | Perennials; short-lived; sexual; caudex not woody. |
Stems | usually 1 per caudex branch, arising from center of rosette near ground surface, (0.8–)2–5 dm, densely pubescent proximally, trichomes short-stalked, 7–12-rayed, 0.1–0.2 mm, densely to sparsely pubescent distally. |
usually 1 per caudex branch, arising from center of rosette near ground surface, 0.7–2.2 dm, glabrous throughout. |
Basal leaves | blade oblanceolate, 3–10 mm wide, margins entire, not ciliate, surfaces densely pubescent, trichomes short-stalked, 7–12-rayed, 0.1–0.2 mm. |
blade oblanceolate, 1–2 mm wide, margins entire, ciliate near petiole base, trichomes (simple), to 0.4 mm, surfaces sparsely to densely pubescent, trichomes short-stalked, 4–8-rayed, 0.03–0.1 mm. |
Cauline leaves | 14–60, concealing stem for most of length; blade auricles 0.5–4 mm, surfaces of distalmost leaves densely pubescent. |
6–11, sometimes concealing stem proximally; blade auricles to 0.5 mm, surfaces of distalmost leaves glabrous. |
Racemes | 20–70-flowered, usually unbranched. |
5–17-flowered, unbranched. |
Flowers | ascending at anthesis; sepals pubescent; petals lavender, 6–9 × 0.8–1.2 mm, glabrous; pollen ellipsoid. |
erect at anthesis; sepals glabrous; petals white, 3–4 × 0.5–0.8 mm, glabrous; pollen ellipsoid. |
Fruiting pedicels | divaricate-ascending, straight, 7–28 mm, pubescent, trichomes appressed, branched. |
erect, straight, 2–4(–7) mm, glabrous. |
Fruits | divaricate-ascending, not appressed to rachis, not secund, usually curved, rarely straight, edges parallel, 4.5–11 cm × 1.5–2 mm; valves glabrous or sparsely pubescent throughout; ovules 140–190 per ovary; style 0.05–0.6 mm. |
erect, usually appressed to rachis, not secund, straight, edges parallel, 2–3.5 cm × 2.2–2.5 mm; valves glabrous; ovules 60–96 per ovary; style 0.05–0.2 mm. |
Seeds | sub-biseriate, 1–1.3 × 0.7–0.8 mm; wing distal or, rarely, absent, 0.05–0.1 mm wide. |
sub-biseriate, 0.7–0.9 × 0.5–0.6 mm, not winged or minutely winged distally. |
Boechera shockleyi |
Boechera williamsii |
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Phenology | Flowering Apr–May. | Flowering May–Jun. |
Habitat | Rock outcrops (primarily dolomite) and gravelly soil in desert scrub, sagebrush, and pinyon-juniper woodlands | Gravelly soil in sagebrush-grassland communities |
Elevation | 1200-2200 m (3900-7200 ft) | 2300-2800 m (7500-9200 ft) |
Distribution |
CA; NV; UT
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WY |
Discussion | Boechera shockleyi is a distinctive sexual species recognizable by the combination of strongly overlapping cauline leaves, a dense covering of minute, 7–12-rayed trichomes, and relatively long, curved, divaricate-ascending fruits with sub-biseriate seeds. It is most similar to B. inyoensis, but differs from that species in its narrower (0.8–1.2 versus 1.2–2 mm) petals, eciliate basal leaves with smaller (0.1–0.2 versus 0.2–0.7 mm) trichomes, greater number (140–190 versus 74–134) of ovules per ovary, and shorter (1–1.3 versus 1.7–2 mm) seeds with distal (versus continuous) wings. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Boechera williamsii is distinguished from all other sexual species of the genus by its combination of relatively short (3–4 mm), white petals, minute, 4–8-rayed trichomes on basal leaves, erect, appressed fruits, and relatively small, wingless seeds. It is known only from Fremont and Sublette counties in west-central Wyoming. The taxon traditionally treated as Arabis (Boechera) williamsii var. saximontana is here recognized as an apomictic species of hybrid origin (see M. D. Windham and I. A. Al-Shehbaz 2007b for detailed comparison). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 406. | FNA vol. 7, p. 411. |
Parent taxa | Brassicaceae > tribe Boechereae > Boechera | Brassicaceae > tribe Boechereae > Boechera |
Sibling taxa | ||
Synonyms | Arabis shockleyi | Arabis williamsii |
Name authority | (Munz) Dorn: Brittonia 55: 3. (2003) | (Rollins) Dorn: Vasc. Pl. Wyoming ed. 3, 376. (2001) |
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