Boechera shockleyi |
Boechera suffrutescens |
|
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Shockley's Rock cress |
woody rock-cress |
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Habit | Perennials; usually short-lived; sexual; caudex not woody, (rarely with persistent, crowded leaf bases). | Perennials; long-lived; sexual or apomictic; caudex woody. |
Stems | usually 1 per caudex branch, arising from center of rosette near ground surface, (0.8–)2–5 dm, densely pubescent proximally, trichomes short-stalked, 7–12-rayed, 0.1–0.2 mm, densely to sparsely pubescent distally. |
usually 1 per caudex branch, arising from center of rosette, elevated above ground surface on woody base, 1–4(–5) dm, usually glabrous proximally, rarely pubescent, trichomes short-stalked, 2-rayed, 0.1–0.3 mm, glabrous distally. |
Basal leaves | blade oblanceolate, 3–10 mm wide, margins entire, not ciliate, surfaces densely pubescent, trichomes short-stalked, 7–12-rayed, 0.1–0.2 mm. |
blade narrowly oblanceolate, 1.5–6 mm wide, margins entire, not ciliate or, rarely, with simple trichomes to 0.7 mm, surfaces glabrous or sparsely to densely pubescent, trichomes short-stalked, 2–6-rayed, 0.07–0.4 mm. |
Cauline leaves | 14–60, concealing stem for most of length; blade auricles 0.5–4 mm, surfaces of distalmost leaves densely pubescent. |
(4–)6–12, sometimes concealing stem proximally; blade auricles 0.5–2 mm, surfaces of distalmost leaves glabrous. |
Racemes | 20–70-flowered, usually unbranched. |
6–12-flowered, usually unbranched. |
Flowers | ascending at anthesis; sepals pubescent; petals lavender, 6–9 × 0.8–1.2 mm, glabrous; pollen ellipsoid. |
pendent at anthesis; sepals glabrous; petals purple or whitish with rose tips, 4.5–6 × 2–2.5 mm, glabrous; pollen ellipsoid or spheroid. |
Fruiting pedicels | divaricate-ascending, straight, 7–28 mm, pubescent, trichomes appressed, branched. |
reflexed, usually abruptly recurved at base, otherwise straight, 4–18 mm, glabrous. |
Fruits | divaricate-ascending, not appressed to rachis, not secund, usually curved, rarely straight, edges parallel, 4.5–11 cm × 1.5–2 mm; valves glabrous or sparsely pubescent throughout; ovules 140–190 per ovary; style 0.05–0.6 mm. |
reflexed to closely pendent, sometimes appressed to rachis, often secund, straight to somewhat curved, edges undulate (not parallel), 3–7(–8) cm × 3–5(–6) mm; valves glabrous; ovules 20–30 per ovary; style 0.4–1.2 mm. |
Seeds | sub-biseriate, 1–1.3 × 0.7–0.8 mm; wing distal or, rarely, absent, 0.05–0.1 mm wide. |
uniseriate, 2.5–5.5 × 1.8–3.5 mm; wing continuous, 0.8–1.5 mm wide. |
Boechera shockleyi |
Boechera suffrutescens |
|
Phenology | Flowering Apr–May. | Flowering Jun–Aug. |
Habitat | Rock outcrops (primarily dolomite) and gravelly soil in desert scrub, sagebrush, and pinyon-juniper woodlands | Rocky slopes and gravelly soil, often with sagebrush |
Elevation | 1200-2200 m (3900-7200 ft) | 1800-3000 m (5900-9800 ft) |
Distribution |
CA; NV; UT
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CA; ID; NV; OR; WA
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Discussion | Boechera shockleyi is a distinctive sexual species recognizable by the combination of strongly overlapping cauline leaves, a dense covering of minute, 7–12-rayed trichomes, and relatively long, curved, divaricate-ascending fruits with sub-biseriate seeds. It is most similar to B. inyoensis, but differs from that species in its narrower (0.8–1.2 versus 1.2–2 mm) petals, eciliate basal leaves with smaller (0.1–0.2 versus 0.2–0.7 mm) trichomes, greater number (140–190 versus 74–134) of ovules per ovary, and shorter (1–1.3 versus 1.7–2 mm) seeds with distal (versus continuous) wings. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Boechera suffrutescens is recognizable by its prominently suffrutescent habit and wide (greater than 3 mm), reflexed fruits. Both sexual and apomictic collections are known; further study is needed to determine whether they truly are conspecific. The taxon previously known as Arabis suffrutescens var. horizontalis appears to be of hybrid origin; it is treated here as a distinct species (see M. D. Windham and I. A. Al-Shehbaz 2007 for detailed comparison). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 406. | FNA vol. 7, p. 409. |
Parent taxa | Brassicaceae > tribe Boechereae > Boechera | Brassicaceae > tribe Boechereae > Boechera |
Sibling taxa | ||
Synonyms | Arabis shockleyi | Arabis suffrutescens, Arabis dianthifolia, Arabis duriuscula |
Name authority | (Munz) Dorn: Brittonia 55: 3. (2003) | (S. Watson) Dorn: Brittonia 55: 3. (2003) |
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