Boechera shockleyi |
Boechera puberula |
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Shockley's Rock cress |
Blue Mountain rockcress, hoary rock-cress, silver rockcress |
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Habit | Perennials; usually short-lived; sexual; caudex not woody, (rarely with persistent, crowded leaf bases). | Perennials; short-lived; sexual; caudex not woody. |
Stems | usually 1 per caudex branch, arising from center of rosette near ground surface, (0.8–)2–5 dm, densely pubescent proximally, trichomes short-stalked, 7–12-rayed, 0.1–0.2 mm, densely to sparsely pubescent distally. |
usually 1 per caudex branch, arising from center of rosette near ground surface, (1–)2–6.3 dm, densely pubescent proximally, trichomes short-stalked, 3–8-rayed, 0.1–0.3(–0.5) mm, similarly pubescent distally. |
Basal leaves | blade oblanceolate, 3–10 mm wide, margins entire, not ciliate, surfaces densely pubescent, trichomes short-stalked, 7–12-rayed, 0.1–0.2 mm. |
blade oblanceolate to linear-oblanceolate, 1.5–5 mm wide, margins usually dentate, not ciliate, surfaces densely pubescent, trichomes short-stalked, 5–12-rayed, 0.05–0.2 mm. |
Cauline leaves | 14–60, concealing stem for most of length; blade auricles 0.5–4 mm, surfaces of distalmost leaves densely pubescent. |
7–45(–65), concealing stem proximally; blade auricles absent or 0.7–3 mm, surfaces of distalmost leaves pubescent. |
Racemes | 20–70-flowered, usually unbranched. |
10–40(–64)-flowered, usually unbranched. |
Flowers | ascending at anthesis; sepals pubescent; petals lavender, 6–9 × 0.8–1.2 mm, glabrous; pollen ellipsoid. |
ascending to descending at anthesis; sepals pubescent; petals white to lavender, 5–9 × 0.8–1.8 mm, glabrous; pollen ellipsoid. |
Fruiting pedicels | divaricate-ascending, straight, 7–28 mm, pubescent, trichomes appressed, branched. |
pendent, recurved distal to horizontal to ascending base, 4–10 mm, pubescent, trichomes appressed, branched. |
Fruits | divaricate-ascending, not appressed to rachis, not secund, usually curved, rarely straight, edges parallel, 4.5–11 cm × 1.5–2 mm; valves glabrous or sparsely pubescent throughout; ovules 140–190 per ovary; style 0.05–0.6 mm. |
closely pendent, rarely appressed to rachis, sometimes somewhat secund, usually straight, edges parallel, 3–6.5 cm × 1.9–2.2 mm; valves pubescent throughout; ovules 38–64 per ovary; style 0.05–0.1 mm. |
Seeds | sub-biseriate, 1–1.3 × 0.7–0.8 mm; wing distal or, rarely, absent, 0.05–0.1 mm wide. |
uniseriate, 1.4–1.8 × 1–1.4 mm; wing continuous, 0.1–0.3 mm wide. |
2n | = 14. |
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Boechera shockleyi |
Boechera puberula |
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Phenology | Flowering Apr–May. | Flowering Apr–Jul. |
Habitat | Rock outcrops (primarily dolomite) and gravelly soil in desert scrub, sagebrush, and pinyon-juniper woodlands | Ledges, rocky slopes, gravelly hillsides in sagebrush, pinyon-juniper, and mountain shrub communities |
Elevation | 1200-2200 m (3900-7200 ft) | 1300-2900 m (4300-9500 ft) |
Distribution |
CA; NV; UT
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CA; NV; OR; UT
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Discussion | Boechera shockleyi is a distinctive sexual species recognizable by the combination of strongly overlapping cauline leaves, a dense covering of minute, 7–12-rayed trichomes, and relatively long, curved, divaricate-ascending fruits with sub-biseriate seeds. It is most similar to B. inyoensis, but differs from that species in its narrower (0.8–1.2 versus 1.2–2 mm) petals, eciliate basal leaves with smaller (0.1–0.2 versus 0.2–0.7 mm) trichomes, greater number (140–190 versus 74–134) of ovules per ovary, and shorter (1–1.3 versus 1.7–2 mm) seeds with distal (versus continuous) wings. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Boechera puberula is a diploid species that appears to intergrade with both B. retrofracta and B. subpinnatifida. The glabrous-fruited specimens discussed by R. C. Rollins (1993) represent apomictic hybrids with other species, primarily B. pendulocarpa. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 406. | FNA vol. 7, p. 399. |
Parent taxa | Brassicaceae > tribe Boechereae > Boechera | Brassicaceae > tribe Boechereae > Boechera |
Sibling taxa | ||
Synonyms | Arabis shockleyi | Arabis puberula, Arabis arida, Arabis beckwithii, Arabis lignipes var. impar, Arabis sabulosa, Arabis subpinnatifida var. beckwithii, Arabis subpinnatifida var. impar, B. beckwithii |
Name authority | (Munz) Dorn: Brittonia 55: 3. (2003) | (Nuttall) Dorn: Brittonia, 55: 3. (2003) |
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