Boechera shockleyi |
Boechera pendulina |
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Shockley's Rock cress |
nodding rock cress, rabbit-ear rockcress |
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Habit | Perennials; usually short-lived; sexual; caudex not woody, (rarely with persistent, crowded leaf bases). | Perennials; short- to long-lived; sexual or apomictic; caudex sometimes woody. |
Stems | usually 1 per caudex branch, arising from center of rosette near ground surface, (0.8–)2–5 dm, densely pubescent proximally, trichomes short-stalked, 7–12-rayed, 0.1–0.2 mm, densely to sparsely pubescent distally. |
usually 2–6 per caudex branch, arising from margin of rosette near ground surface, or arising laterally proximal to sterile shoots, 0.6–3(–3.7) dm, sparsely to densely pubescent proximally, trichomes simple, 0.3–0.8 mm, glabrous distally. |
Basal leaves | blade oblanceolate, 3–10 mm wide, margins entire, not ciliate, surfaces densely pubescent, trichomes short-stalked, 7–12-rayed, 0.1–0.2 mm. |
blade oblanceolate to obovate, 1.5–6 mm wide, margins usually entire, rarely dentate, ciliate throughout, trichomes (usually simple), 0.4–1 mm, surfaces pubescent, trichomes simple and short- and long-stalked, 2-rayed, 0.3–0.8 mm. |
Cauline leaves | 14–60, concealing stem for most of length; blade auricles 0.5–4 mm, surfaces of distalmost leaves densely pubescent. |
2–10(–13), not concealing stem; blade auricles absent or, rarely, to 0.7 mm, surfaces of distalmost leaves glabrous or margins sparsely ciliate. |
Racemes | 20–70-flowered, usually unbranched. |
4–14-flowered, usually unbranched. |
Flowers | ascending at anthesis; sepals pubescent; petals lavender, 6–9 × 0.8–1.2 mm, glabrous; pollen ellipsoid. |
divaricate-ascending at anthesis; sepals usually sparsely pubescent, rarely glabrous; petals whitish to pale lavender, 4–6 × 1–1.5 mm, glabrous; pollen ellipsoid or spheroid. |
Fruiting pedicels | divaricate-ascending, straight, 7–28 mm, pubescent, trichomes appressed, branched. |
divaricate-ascending to horizontal, curved or angled downward, 3–7(–10) mm, usually glabrous, rarely with some simple trichomes. |
Fruits | divaricate-ascending, not appressed to rachis, not secund, usually curved, rarely straight, edges parallel, 4.5–11 cm × 1.5–2 mm; valves glabrous or sparsely pubescent throughout; ovules 140–190 per ovary; style 0.05–0.6 mm. |
widely pendent, not appressed to rachis, not secund, curved to nearly straight, edges parallel, 2.2–4 cm × 1.2–2.1 mm; valves glabrous; ovules 40–70(–90) per ovary; style 0.1–0.3(–0.5) mm. |
Seeds | sub-biseriate, 1–1.3 × 0.7–0.8 mm; wing distal or, rarely, absent, 0.05–0.1 mm wide. |
biseriate, 0.9–1.2 × 0.6–0.9 mm, usually not winged. |
2n | = 14, 21. |
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Boechera shockleyi |
Boechera pendulina |
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Phenology | Flowering Apr–May. | Flowering Apr–Jun. |
Habitat | Rock outcrops (primarily dolomite) and gravelly soil in desert scrub, sagebrush, and pinyon-juniper woodlands | Rock outcrops, open gravelly flats and hillsides in sagebrush, pinyon-juniper, mountain mahogany, open conifer forests |
Elevation | 1200-2200 m (3900-7200 ft) | 1600-3100 m (5200-10200 ft) |
Distribution |
CA; NV; UT
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AZ; CA; CO; NV; UT; WY
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Discussion | Boechera shockleyi is a distinctive sexual species recognizable by the combination of strongly overlapping cauline leaves, a dense covering of minute, 7–12-rayed trichomes, and relatively long, curved, divaricate-ascending fruits with sub-biseriate seeds. It is most similar to B. inyoensis, but differs from that species in its narrower (0.8–1.2 versus 1.2–2 mm) petals, eciliate basal leaves with smaller (0.1–0.2 versus 0.2–0.7 mm) trichomes, greater number (140–190 versus 74–134) of ovules per ovary, and shorter (1–1.3 versus 1.7–2 mm) seeds with distal (versus continuous) wings. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Although included in Arabis (Boechera) demissa by some authors (e.g., S. L. Welsh et al. 2003; N. H. Holmgren 2005b), B. pendulina is readily distinguished from that species (see M. D. Windham and I. A. Al-Shehbaz 2006 for detailed comparison). Typical collections are sexual diploids, whereas the type of var. russeola is a triploid apomict; further study is needed to determine if the two are conspecific. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 406. | FNA vol. 7, p. 395. |
Parent taxa | Brassicaceae > tribe Boechereae > Boechera | Brassicaceae > tribe Boechereae > Boechera |
Sibling taxa | ||
Synonyms | Arabis shockleyi | Arabis pendulina, Arabis demissa var. russeola, Arabis diehlii, Arabis pendulina var. russeola, Arabis setulosa, B. demissa var. pendulina, B. demissa var. russeola, B. pendulina var. russeola |
Name authority | (Munz) Dorn: Brittonia 55: 3. (2003) | (Greene) W. A. Weber: Phytologia 51: 370. (1982) |
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