Boechera shockleyi |
Boechera lemmonii |
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Shockley's Rock cress |
Lemmon's rock-cress, Lemmon's suncress, wind-blown suncress |
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Habit | Perennials; usually short-lived; sexual; caudex not woody, (rarely with persistent, crowded leaf bases). | Perennials; long-lived; (somewhat cespitose); sexual or apomictic; caudex woody. |
Stems | usually 1 per caudex branch, arising from center of rosette near ground surface, (0.8–)2–5 dm, densely pubescent proximally, trichomes short-stalked, 7–12-rayed, 0.1–0.2 mm, densely to sparsely pubescent distally. |
usually 1 per caudex branch, arising from center of rosette near ground surface, or arising laterally proximal to sterile shoots, 0.5–2(–2.5) dm, glabrous or sparsely pubescent proximally, trichomes short-stalked, 2–6-rayed, 0.1–0.2 mm, glabrous distally. |
Basal leaves | blade oblanceolate, 3–10 mm wide, margins entire, not ciliate, surfaces densely pubescent, trichomes short-stalked, 7–12-rayed, 0.1–0.2 mm. |
blade oblanceolate to obovate, 1.5–5 mm wide, margins usually entire, rarely slightly dentate, ciliate along petiole, surfaces densely to sparsely pubescent, trichomes short-stalked, 3–9-rayed, 0.1–0.2 mm. |
Cauline leaves | 14–60, concealing stem for most of length; blade auricles 0.5–4 mm, surfaces of distalmost leaves densely pubescent. |
2–8(–12), not concealing stem; blade auricles absent or 0.1–0.5 mm, surfaces of distalmost leaves glabrous or sparsely pubescent. |
Racemes | 20–70-flowered, usually unbranched. |
3–12(–17)-flowered, usually unbranched. |
Flowers | ascending at anthesis; sepals pubescent; petals lavender, 6–9 × 0.8–1.2 mm, glabrous; pollen ellipsoid. |
divaricate-ascending at anthesis; sepals glabrous or sparsely pubescent; petals purple to lavender, 3.5–6 × 1–1.5 mm; pollen ellipsoid or spheroid. |
Fruiting pedicels | divaricate-ascending, straight, 7–28 mm, pubescent, trichomes appressed, branched. |
divaricate-ascending to slightly descending, usually slightly recurved, rarely straight, 2–6 mm, glabrous or sparsely pubescent, trichomes appressed, branched. |
Fruits | divaricate-ascending, not appressed to rachis, not secund, usually curved, rarely straight, edges parallel, 4.5–11 cm × 1.5–2 mm; valves glabrous or sparsely pubescent throughout; ovules 140–190 per ovary; style 0.05–0.6 mm. |
divaricate-ascending to slightly descending, not appressed to rachis, secund, straight or curved, edges parallel, (1.6–) 2–4.4 cm × 1.6–2.3 mm; valves glabrous; ovules 28 40(–44) per ovary; style 0.1–0.2 mm. |
Seeds | sub-biseriate, 1–1.3 × 0.7–0.8 mm; wing distal or, rarely, absent, 0.05–0.1 mm wide. |
uniseriate, 1.3–2 × 1–1.5 mm; wing continuous, 0.1–0.5 mm wide. |
2n | = 14. |
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Boechera shockleyi |
Boechera lemmonii |
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Phenology | Flowering Apr–May. | Flowering Jun–Aug. |
Habitat | Rock outcrops (primarily dolomite) and gravelly soil in desert scrub, sagebrush, and pinyon-juniper woodlands | Cliffs, talus slopes, and gravelly soil in alpine and subalpine habitats |
Elevation | 1200-2200 m (3900-7200 ft) | 2100-4400 m (6900-14400 ft) |
Distribution |
CA; NV; UT
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AK; CA; CO; ID; MT; NV; OR; UT; WA; WY; AB; BC; YT
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Discussion | Boechera shockleyi is a distinctive sexual species recognizable by the combination of strongly overlapping cauline leaves, a dense covering of minute, 7–12-rayed trichomes, and relatively long, curved, divaricate-ascending fruits with sub-biseriate seeds. It is most similar to B. inyoensis, but differs from that species in its narrower (0.8–1.2 versus 1.2–2 mm) petals, eciliate basal leaves with smaller (0.1–0.2 versus 0.2–0.7 mm) trichomes, greater number (140–190 versus 74–134) of ovules per ovary, and shorter (1–1.3 versus 1.7–2 mm) seeds with distal (versus continuous) wings. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Boechera lemmonii is easily recognized by its combination of secund fruits, mat-forming habit, purplish sepals, and obovate-oblanceolate cauline leaves. Both sexual and apomictic collections are known; further study is needed to determine whether they truly are conspecific. The taxa traditionally treated as Arabis (Boechera) lemmonii vars. depauperata, drepanoloba, and paddoensis are apomictic hybrids here recognized as distinct species (see M. D. Windham and I. A. Al-Shehbaz 2007 for detailed comparison). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 406. | FNA vol. 7, p. 388. |
Parent taxa | Brassicaceae > tribe Boechereae > Boechera | Brassicaceae > tribe Boechereae > Boechera |
Sibling taxa | ||
Synonyms | Arabis shockleyi | Arabis lemmonii, Arabis bracteolata, Arabis canescens var. latifolia, Arabis codyi, Arabis egglestonii, Arabis kennedyi, Arabis latifolia, Arabis oreocallis, Arabis polyclada, Arabis semisepulta |
Name authority | (Munz) Dorn: Brittonia 55: 3. (2003) | (S. Watson) W. A. Weber: Phytologia 51: 370. (1982) |
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