Boechera saximontana |
Boechera platysperma |
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broad-seed rock-cress, flat-seed rock-cress, pioneer rock-cress |
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Habit | Perennials; short-lived; apomictic; caudex usually not woody. | Perennials; long-lived; sexual; caudex woody. |
Stems | usually 1 per caudex branch, arising from center of rosette near ground surface, 0.7–3 dm, densely pubescent proximally, trichomes short-stalked, 2–6-rayed, 0.06–0.25 mm, glabrous distally. |
usually 1 per caudex branch, arising from center of rosette, elevated on woody base above or near ground surface, 0.6–2.8(–3.5) dm, densely pubescent proximally, trichomes short-stalked, 2–5-rayed, 0.1–0.3 mm, glabrous or sparsely pubescent distally. |
Basal leaves | blade narrowly oblanceolate, 1–3 mm wide, margins entire, ciliate near petiole base, trichomes (simple and spurred), to 0.4 mm, surfaces sparsely to densely pubescent, trichomes short-stalked, 4–8-rayed, 0.04–0.15 mm. |
blade narrowly oblanceolate, 3–7(–10) mm wide, margins entire, ciliate near petiole base, trichomes (simple or spurred), to 1 mm, surfaces densely pubescent, trichomes short-stalked, 2–4 (or 5)-rayed 0.1–0.3 mm. |
Cauline leaves | 5–9, rarely concealing stem proximally; blade auricles to 0.3–1 mm, surfaces of distalmost leaves glabrous. |
3–7(–12), not concealing stem; blade auricles absent, surfaces of distalmost leaves glabrous or sparsely pubescent. |
Racemes | 5–13-flowered, usually unbranched. |
2–7-flowered, unbranched. |
Flowers | ascending at anthesis; sepals pubescent; petals lavender, 4–5 × 0.5–0.8 mm, glabrous; pollen spheroid. |
ascending at anthesis; sepals pubescent; petals creamy white to purplish, 4–6 × 1–2 mm, sparsely pubescent (often some trichomes abaxially); pollen ellipsoid. |
Fruiting pedicels | divaricate-ascending, straight, 3–8 mm, glabrous. |
ascending, straight, 3–13 mm, glabrous or sparsely pubescent, trichomes appressed, branched. |
Fruits | divaricate-ascending, not appressed to rachis, not secund, straight, edges parallel, (2.5–)3–4.7 cm × 1.2–1.7 mm; valves glabrous; ovules 78–112 per ovary; style 0.1–0.5 mm. |
ascending, not appressed to rachis, not secund, straight to slightly curved, edges undulate (not parallel), (2.5–)4–8.5 cm × 3–5.5 mm; valves glabrous; ovules 16–44 per ovary; style 0.05–1 mm. |
Seeds | uniseriate to sub-biseriate, 1.1–1.4 × 0.7–0.9 mm; wing continuous, 0.05–0.12 mm wide. |
uniseriate, 3–6(–8) × 2–3.5 mm; wing continuous, 0.8–2.5 mm wide. |
2n | = 14. |
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Boechera saximontana |
Boechera platysperma |
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Phenology | Flowering Jun–Jul. | Flowering Jun–Aug. |
Habitat | Rocky soil in sagebrush and open conifer forests | Rock outcrops and gravelly soil in dry pine forests and lodgepole-chaparral woodlands |
Elevation | 2400-2900 m (7900-9500 ft) | 1600-3000 m (5200-9800 ft) |
Distribution |
ID; WY |
CA; OR
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Discussion | Boechera saximontana is an apomictic taxon that has been treated as a variety of either Arabis microphylla (R. C. Rollins 1941) or A. (Boechera) williamsii (Rollins 1993; R. D. Dorn 2001). It is easily distinguished from typical collections of those species (see M. D. Windham and I. A. Al-Shehbaz 2007b for detailed comparison), but it is likely that one (or both) were involved in its hybrid origin. It is known from Blaine, Custer, and Lemhi counties in Idaho, and Big Horn and Fremont counties in Wyoming. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Previous authors have adopted broader circumscriptions of Boechera platysperma, including plants herein called B. covillei, B. elkoensis, and B. howellii. In our opinion, the differences between these taxa are sufficient to justify their treatment as distinct species (see M. D. Windham and I. A. Al-Shehbaz 2006, 2007, 2007b for detailed comparison). Reports of B. platysperma from Nevada (e.g., R. C. Rollins 1993; N. H. Holmgren 2005b) are based on plants of B. howellii or apomictic hybrids with other species. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 404. | FNA vol. 7, p. 397. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Arabis microphylla var. saximontana, Arabis pendulocarpa var. saximontana, Arabis williamsii var. saximontana, B. williamsii var. saximontana | Arabis platysperma, Arabis inamoena, Arabis oligantha |
Name authority | (Rollins) Windham & Al-Shehbaz: Harvard Pap. Bot. 12: 248. (2007) | (A. Gray) Al-Shehbaz: Novon 13: 388. (2003) |
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