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broad-seed rock-cress, flat-seed rock-cress, pioneer rock-cress

Habit Perennials; short-lived; apomictic; caudex usually not woody. Perennials; long-lived; sexual; caudex woody.
Stems

usually 1 per caudex branch, arising from center of rosette near ground surface, 0.7–3 dm, densely pubescent proximally, trichomes short-stalked, 2–6-rayed, 0.06–0.25 mm, glabrous distally.

usually 1 per caudex branch, arising from center of rosette, elevated on woody base above or near ground surface, 0.6–2.8(–3.5) dm, densely pubescent proximally, trichomes short-stalked, 2–5-rayed, 0.1–0.3 mm, glabrous or sparsely pubescent distally.

Basal leaves

blade narrowly oblanceolate, 1–3 mm wide, margins entire, ciliate near petiole base, trichomes (simple and spurred), to 0.4 mm, surfaces sparsely to densely pubescent, trichomes short-stalked, 4–8-rayed, 0.04–0.15 mm.

blade narrowly oblanceolate, 3–7(–10) mm wide, margins entire, ciliate near petiole base, trichomes (simple or spurred), to 1 mm, surfaces densely pubescent, trichomes short-stalked, 2–4 (or 5)-rayed 0.1–0.3 mm.

Cauline leaves

5–9, rarely concealing stem proximally;

blade auricles to 0.3–1 mm, surfaces of distalmost leaves glabrous.

3–7(–12), not concealing stem;

blade auricles absent, surfaces of distalmost leaves glabrous or sparsely pubescent.

Racemes

5–13-flowered, usually unbranched.

2–7-flowered, unbranched.

Flowers

ascending at anthesis;

sepals pubescent;

petals lavender, 4–5 × 0.5–0.8 mm, glabrous;

pollen spheroid.

ascending at anthesis;

sepals pubescent;

petals creamy white to purplish, 4–6 × 1–2 mm, sparsely pubescent (often some trichomes abaxially);

pollen ellipsoid.

Fruiting pedicels

divaricate-ascending, straight, 3–8 mm, glabrous.

ascending, straight, 3–13 mm, glabrous or sparsely pubescent, trichomes appressed, branched.

Fruits

divaricate-ascending, not appressed to rachis, not secund, straight, edges parallel, (2.5–)3–4.7 cm × 1.2–1.7 mm;

valves glabrous;

ovules 78–112 per ovary;

style 0.1–0.5 mm.

ascending, not appressed to rachis, not secund, straight to slightly curved, edges undulate (not parallel), (2.5–)4–8.5 cm × 3–5.5 mm;

valves glabrous;

ovules 16–44 per ovary;

style 0.05–1 mm.

Seeds

uniseriate to sub-biseriate, 1.1–1.4 × 0.7–0.9 mm;

wing continuous, 0.05–0.12 mm wide.

uniseriate, 3–6(–8) × 2–3.5 mm;

wing continuous, 0.8–2.5 mm wide.

2n

= 14.

Boechera saximontana

Boechera platysperma

Phenology Flowering Jun–Jul. Flowering Jun–Aug.
Habitat Rocky soil in sagebrush and open conifer forests Rock outcrops and gravelly soil in dry pine forests and lodgepole-chaparral woodlands
Elevation 2400-2900 m (7900-9500 ft) 1600-3000 m (5200-9800 ft)
Distribution
from FNA
ID; WY
[BONAP county map]
from FNA
CA; OR
[WildflowerSearch map]
[BONAP county map]
Discussion

Boechera saximontana is an apomictic taxon that has been treated as a variety of either Arabis microphylla (R. C. Rollins 1941) or A. (Boechera) williamsii (Rollins 1993; R. D. Dorn 2001). It is easily distinguished from typical collections of those species (see M. D. Windham and I. A. Al-Shehbaz 2007b for detailed comparison), but it is likely that one (or both) were involved in its hybrid origin. It is known from Blaine, Custer, and Lemhi counties in Idaho, and Big Horn and Fremont counties in Wyoming.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Previous authors have adopted broader circumscriptions of Boechera platysperma, including plants herein called B. covillei, B. elkoensis, and B. howellii. In our opinion, the differences between these taxa are sufficient to justify their treatment as distinct species (see M. D. Windham and I. A. Al-Shehbaz 2006, 2007, 2007b for detailed comparison). Reports of B. platysperma from Nevada (e.g., R. C. Rollins 1993; N. H. Holmgren 2005b) are based on plants of B. howellii or apomictic hybrids with other species.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 404. FNA vol. 7, p. 397.
Parent taxa Brassicaceae > tribe Boechereae > Boechera Brassicaceae > tribe Boechereae > Boechera
Sibling taxa
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
Synonyms Arabis microphylla var. saximontana, Arabis pendulocarpa var. saximontana, Arabis williamsii var. saximontana, B. williamsii var. saximontana Arabis platysperma, Arabis inamoena, Arabis oligantha
Name authority (Rollins) Windham & Al-Shehbaz: Harvard Pap. Bot. 12: 248. (2007) (A. Gray) Al-Shehbaz: Novon 13: 388. (2003)
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