Boechera saximontana |
Boechera johnstonii |
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Johnston's rockcress |
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Habit | Perennials; short-lived; apomictic; caudex usually not woody. | Perennials; long-lived; (cespitose); sexual; caudex woody. |
Stems | usually 1 per caudex branch, arising from center of rosette near ground surface, 0.7–3 dm, densely pubescent proximally, trichomes short-stalked, 2–6-rayed, 0.06–0.25 mm, glabrous distally. |
usually 1 per caudex branch, arising from center of rosette near ground surface, 0.5–2 dm, densely pubescent proximally, trichomes short-stalked, 4–10-rayed, 0.07–0.15 mm, sparsely to densely pubescent distally. |
Basal leaves | blade narrowly oblanceolate, 1–3 mm wide, margins entire, ciliate near petiole base, trichomes (simple and spurred), to 0.4 mm, surfaces sparsely to densely pubescent, trichomes short-stalked, 4–8-rayed, 0.04–0.15 mm. |
blade narrowly oblanceolate, 1.5–4 mm wide, margins entire, not ciliate, surfaces densely pubescent, trichomes short-stalked, 6–14-rayed, 0.07–0.15 mm. |
Cauline leaves | 5–9, rarely concealing stem proximally; blade auricles to 0.3–1 mm, surfaces of distalmost leaves glabrous. |
4–10, often concealing stem proximally; blade auricles absent, surfaces of distalmost leaves densely pubescent. |
Racemes | 5–13-flowered, usually unbranched. |
10–18-flowered, unbranched. |
Flowers | ascending at anthesis; sepals pubescent; petals lavender, 4–5 × 0.5–0.8 mm, glabrous; pollen spheroid. |
ascending at anthesis; sepals pubescent; petals purple, 9–14 × 2–4 mm, glabrous; pollen ellipsoid. |
Fruiting pedicels | divaricate-ascending, straight, 3–8 mm, glabrous. |
divaricate-ascending, straight, 5–14 mm, pubescent, trichomes appressed, branched. |
Fruits | divaricate-ascending, not appressed to rachis, not secund, straight, edges parallel, (2.5–)3–4.7 cm × 1.2–1.7 mm; valves glabrous; ovules 78–112 per ovary; style 0.1–0.5 mm. |
divaricate-ascending, not appressed to rachis, not secund, straight, edges parallel, 4–6 cm × 2.5–4 mm; valves glabrous; ovules 26–34 per ovary; style (0.7–)1–2 mm. |
Seeds | uniseriate to sub-biseriate, 1.1–1.4 × 0.7–0.9 mm; wing continuous, 0.05–0.12 mm wide. |
uniseriate, 1.9–2.7 × 1.5–2.2 mm; wing continuous, 0.3–0.7 mm wide. |
Boechera saximontana |
Boechera johnstonii |
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Phenology | Flowering Jun–Jul. | Flowering Feb–Mar. |
Habitat | Rocky soil in sagebrush and open conifer forests | Rocky areas and gravelly soil in chaparral and oak-pine savannas |
Elevation | 2400-2900 m (7900-9500 ft) | 1300-1700 m (4300-5600 ft) |
Distribution |
ID; WY |
CA |
Discussion | Boechera saximontana is an apomictic taxon that has been treated as a variety of either Arabis microphylla (R. C. Rollins 1941) or A. (Boechera) williamsii (Rollins 1993; R. D. Dorn 2001). It is easily distinguished from typical collections of those species (see M. D. Windham and I. A. Al-Shehbaz 2007b for detailed comparison), but it is likely that one (or both) were involved in its hybrid origin. It is known from Blaine, Custer, and Lemhi counties in Idaho, and Big Horn and Fremont counties in Wyoming. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Boechera johnstonii and B. hirshbergiae are too similar to be considered distinct species. Even with the expanded circumscription, this distinctive sexual species is known only from the area of Cuyamaca Lake (San Diego County) and the San Jacinto Mountains (Riverside County). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 404. | FNA vol. 7, p. 386. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Arabis microphylla var. saximontana, Arabis pendulocarpa var. saximontana, Arabis williamsii var. saximontana, B. williamsii var. saximontana | Arabis johnstonii, Arabis hirshbergiae, B. hirshbergiae |
Name authority | (Rollins) Windham & Al-Shehbaz: Harvard Pap. Bot. 12: 248. (2007) | (Munz) Al-Shehbaz: Novon 13: 386. (2003) |
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