Boechera saximontana |
Boechera holboellii |
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Habit | Perennials; short-lived; apomictic; caudex usually not woody. | Biennials or perennials; short-lived; apomictic or sexual; caudex present or absent. |
Stems | usually 1 per caudex branch, arising from center of rosette near ground surface, 0.7–3 dm, densely pubescent proximally, trichomes short-stalked, 2–6-rayed, 0.06–0.25 mm, glabrous distally. |
usually 1 per caudex branch, arising from center of rosette near ground surface, 2–6.7 dm, densely pubescent proximally, trichomes short-stalked, 4–8-rayed, 0.1–0.3 mm, glabrous or sparsely pubescent distally. |
Basal leaves | blade narrowly oblanceolate, 1–3 mm wide, margins entire, ciliate near petiole base, trichomes (simple and spurred), to 0.4 mm, surfaces sparsely to densely pubescent, trichomes short-stalked, 4–8-rayed, 0.04–0.15 mm. |
blade oblanceolate, 2–5 mm wide, margins entire, sometimes ciliate near petiole base, trichomes (usually simple), to 0.7 mm, surfaces sparsely to moderately pubescent, trichomes short-stalked, 4–8-rayed, 0.1–0.3 mm. |
Cauline leaves | 5–9, rarely concealing stem proximally; blade auricles to 0.3–1 mm, surfaces of distalmost leaves glabrous. |
7–40, often concealing stem proximally; blade auricles 1–4 mm, surfaces of distalmost leaves glabrous or sparsely pubescent. |
Racemes | 5–13-flowered, usually unbranched. |
10–60-flowered, usually unbranched. |
Flowers | ascending at anthesis; sepals pubescent; petals lavender, 4–5 × 0.5–0.8 mm, glabrous; pollen spheroid. |
divaricate to pendent at anthesis; sepals pubescent; petals white, 4–8 × 1–2 mm, glabrous; pollen ellipsoid or spheroid. |
Fruiting pedicels | divaricate-ascending, straight, 3–8 mm, glabrous. |
descending to reflexed, most abruptly recurved near base, 5–10 mm, glabrous or sparsely pubescent, trichomes appressed, branched. |
Fruits | divaricate-ascending, not appressed to rachis, not secund, straight, edges parallel, (2.5–)3–4.7 cm × 1.2–1.7 mm; valves glabrous; ovules 78–112 per ovary; style 0.1–0.5 mm. |
descending to reflexed, not appressed to rachis, strongly secund, often curved, edges parallel, 0.35–0.65 cm × (2–)2.2–2.5 mm; valves glabrous; ovules 80–102 per ovary. |
Seeds | uniseriate to sub-biseriate, 1.1–1.4 × 0.7–0.9 mm; wing continuous, 0.05–0.12 mm wide. |
uniseriate to sub-biseriate, 1.4–1.7 × 1–1.2 mm; wing continuous, to 0.2 mm wide. |
2n | = 14, 21. |
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Boechera saximontana |
Boechera holboellii |
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Phenology | Flowering Jun–Jul. | Flowering Jul–Aug. |
Habitat | Rocky soil in sagebrush and open conifer forests | Rocky slopes and gravelly soil near coast |
Elevation | 2400-2900 m (7900-9500 ft) | 100-300 m (300-1000 ft) |
Distribution |
ID; WY |
Greenland |
Discussion | Boechera saximontana is an apomictic taxon that has been treated as a variety of either Arabis microphylla (R. C. Rollins 1941) or A. (Boechera) williamsii (Rollins 1993; R. D. Dorn 2001). It is easily distinguished from typical collections of those species (see M. D. Windham and I. A. Al-Shehbaz 2007b for detailed comparison), but it is likely that one (or both) were involved in its hybrid origin. It is known from Blaine, Custer, and Lemhi counties in Idaho, and Big Horn and Fremont counties in Wyoming. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Both sexual diploid and apomictic triploid collections of Boechera holboellii are known; available data suggest that the triploid is an autopolyploid derivative of the sexual diploid but further study is needed. In the narrow circumscription adopted here, it appears to be restricted to Greenland (see discussion of B. quebecensis in M. D. Windham and I. A. Al-Shehbaz 2007b). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 404. | FNA vol. 7, p. 384. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Arabis microphylla var. saximontana, Arabis pendulocarpa var. saximontana, Arabis williamsii var. saximontana, B. williamsii var. saximontana | Arabis holboellii, Arabis holboellii var. tenuis, B. tenuis |
Name authority | (Rollins) Windham & Al-Shehbaz: Harvard Pap. Bot. 12: 248. (2007) | (Hornemann) Á. Löve & D. Löve: Bot. Not. 128: 513. (1976) |
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