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Graham's rockcress, Graham's suncress

Habit Perennials; long-lived; sexual; caudex woody. Biennials or perennials; short-lived; apomictic; caudex present or absent.
Stems

usually 3–5 per caudex branch, arising from margin of rosette, elevated above ground surface on woody base, 1–2.5 dm, densely pubescent proximally, trichomes short-stalked, 2–4 (or 5)-rayed, 0.2–0.4 mm, moderately to sparsely pubescent distally.

usually 1 per caudex branch, arising from center of rosette near ground surface, (1.2–)2–12 dm, glabrous or pubescent proximally, trichomes simple and sessile or subsessile, 2 or 3(–6)-rayed, to 1 mm, glabrous distally.

Basal leaves

blade oblanceolate, 3–5 mm wide, margins entire, ciliate proximally, trichomes (simple), to 0.6 mm, surfaces densely pubescent, trichomes short-stalked, 2–4 (or 5)-rayed, 0.15–0.4 mm.

blade oblanceolate, 1.5–10(–20) mm wide, margins entire or dentate, ciliate proximally, trichomes (simple), to 1 mm, surfaces sparsely to densely pubescent, trichomes sessile or subsessile, 2–4(–7)-rayed, 0.2–0.6 mm.

Cauline leaves

5–9, not concealing stem;

blade auricles 0.5–1 mm, surfaces of distalmost leaves sparsely to moderately pubescent.

(10–)13–52, often concealing stem proximally;

blade auricles 1–5 mm, surfaces of distalmost leaves glabrous.

Racemes

5–11-flowered, usually unbranched.

16–88-flowered, usually unbranched.

Flowers

divaricate-ascending at anthesis;

sepals pubescent;

petals pale purple, 6–8 × 1.5–2 mm, glabrous;

pollen ellipsoid.

divaricate at anthesis;

sepals glabrous or pubescent;

petals usually white, rarely lavender, 5.5–8 × 1–2 mm, glabrous;

pollen spheroid.

Fruiting pedicels

descending, straight to slightly recurved, 4–6 mm, pubescent, trichomes appressed, branched.

divaricate-ascending to descending, usually gently curved downward, rarely straight, 5–22(–30) mm, glabrous.

Fruits

(immature) pendent, not appressed to rachis, not secund, straight, edges parallel;

valves glabrous.

divaricate-ascending to pendent, not appressed to rachis, not secund, straight or slightly curved, edges parallel, 3.5–9 cm × 1–1.8 mm;

valves glabrous;

ovules 84–146 per ovary;

style 0.05–0.5 mm.

Seeds

not seen.

uniseriate to sub-biseriate, 1–1.8 × 0.8–1.2 mm;

wing continuous, 0.1–0.25 mm wide.

2n

= 21.

Boechera rollinsiorum

Boechera grahamii

Phenology Flowering Jun. Flowering May–Aug.
Habitat Metamorphosed igneous gravel on steep slopes Rocky slopes and sandy soil in prairies and open forests
Elevation ca. 2100 m (ca. 6900 ft) 100-3200 m (300-10500 ft)
Distribution
from FNA
ID
[BONAP county map]
from FNA
AK; CO; MI; MN; MT; ND; NY; SD; VT; WA; WI; WY; AB; BC; MB; NT; ON; QC; SK; YT
[WildflowerSearch map]
[BONAP county map]
Discussion

Boechera rollinsiorum shows superficial similarities to both B. glareosa and B. lasiocarpa (see M. D. Windham and I. A. Al-Shehbaz 2006 for detailed comparison). It is known thus far only from the type collection below Galena Summit in central Idaho.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Morphological evidence suggests that Boechera grahamii is an apomictic species that arose through hybridization between B. collinsii and B. stricta. Previous authors have assigned these specimens to Arabis (Boechera) divaricarpa (see M. D. Windham and I. A. Al-Shehbaz 2007b for detailed comparison); if these taxa are treated as conspecific, the name B. grahamii has priority.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 403. FNA vol. 7, p. 382.
Parent taxa Brassicaceae > tribe Boechereae > Boechera Brassicaceae > tribe Boechereae > Boechera
Sibling taxa
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollei, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
Synonyms Turritis grahamii, Arabis boivinii, Arabis bourgovii, Arabis confinis, Arabis confinis var. brachycarpa, Arabis dacotica, Arabis divaricarpa var. dacotica, Arabis divaricarpa var. hemicylindrica, Arabis divaricarpa var. stenocarpa, Arabis drummondii var. brachycarpa, Arabis holboellii var. brachycarpa, B. brachycarpa, Turritis brachycarpa
Name authority Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 81. (2006) (Lehmann) Windham & Al-Shehbaz: Harvard Pap. Bot. 12: 241. (2007)
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