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Rolle's rock-cress

Tulare County rock cress

Habit Perennials; long-lived; sexual; caudex woody. Perennials; usually long-lived; sexual; caudex woody, (often with persistent, crowded leaf bases).
Stems

usually 1 per caudex branch, arising from center of rosette, elevated above ground surface on woody base, 1.5–2.5 dm, glabrous throughout.

usually 2–5 per caudex branch, arising from margin of rosette near ground surface, or arising laterally proximal to sterile shoots, 0.2–0.8 dm, pubescent proximally, trichomes simple and short-stalked, 2- or 3-rayed, to 0.4 mm, glabrescent distally.

Basal leaves

blade oblanceolate, 3–8 mm wide, margins entire, ciliate proximally, trichomes (simple), 0.2–0.7 mm, surfaces glabrous or sparsely pubescent, trichomes short-stalked, 2–4-rayed, 0.2–0.4 mm.

blade linear, 0.8–1.5 mm wide, margins entire, ciliate, trichomes (simple and spurred), to 0.8 mm, surfaces moderately pubescent, trichomes short-stalked, 2–4-rayed, 0.05–0.4 mm.

Cauline leaves

6–12, sometimes concealing stem proximally;

blade auricles 0.5–2.5 mm, surfaces of distalmost leaves glabrous.

2–4, not concealing stem;

blade auricles absent, surfaces of distalmost leaves pubescent.

Racemes

3–7-flowered, unbranched.

2–5-flowered, unbranched.

Flowers

divaricate-ascending at anthesis;

sepals glabrous;

petals creamy white, 8–11 × 2–2.5 mm, glabrous;

pollen ellipsoid.

erect at anthesis;

sepals pubescent;

petals white, 3.5–5 × 0.7–1 mm, glabrous;

pollen ellipsoid.

Fruiting pedicels

strongly curved proximally, 4–6 mm, glabrous.

erect to ascending, straight, 2–7 mm, usually pubescent, rarely glabrous, trichomes subappressed, branched.

Fruits

pendent, not appressed to rachis, not secund, straight to slightly curved, edges undulate (not parallel), 3.5–6 cm × 2.5–3.5 mm;

valves glabrous;

ovules 14–22 per ovary;

style 0.5–1 mm.

erect to ascending, often appressed to rachis, not secund, straight, edges parallel, (1.3–)2–3.3 cm × 4–5 mm;

valves glabrous;

ovules 8–12 per ovary;

style 0.05–0.4 mm.

Seeds

uniseriate, 3–4 × 1.5–2 mm;

wing distal and proximal, 0.3–0.6 mm wide.

uniseriate, 3–5 × 2.5–4.5 mm;

wing continuous, 0.8–2 mm wide distally.

Boechera rollei

Boechera pygmaea

Phenology Flowering Aug. Flowering May–Jul.
Habitat Peridotite rocks on sparsely forested slopes Barren flats of arkosic gravel
Elevation 1600-1800 m (5200-5900 ft) 2400-3200 m (7900-10500 ft)
Distribution
from FNA
CA
[BONAP county map]
from FNA
CA
[BONAP county map]
Discussion

Boechera rollei is easily distinguished from the habitally similar B. suffrutescens by its longer (8–11 versus 4.5–6 mm) petals, narrower (2.5–3.5 versus 3–6 mm) fruits, and narrower (0.3–0.6 versus 0.8–1.5 mm) seed wings. This distinctive sexual species is known only from the vicinity of the type locality on the divide between the Applegate and Klamath Rivers in Siskiyou County.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Despite the distinctive morphology of Boechera pygmaea, there is evidence that it is capable of producing fertile hybrids with B. stricta. It is known only from Inyo and Tulare counties in the southern Sierra Nevada.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 403. FNA vol. 7, p. 401.
Parent taxa Brassicaceae > tribe Boechereae > Boechera Brassicaceae > tribe Boechereae > Boechera
Sibling taxa
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. quebecensis, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
Synonyms Arabis rollei Arabis pygmaea
Name authority (Rollins) Al-Shehbaz: Novon 13: 389. (2003) (Rollins) Al-Shehbaz: Novon 13: 388. (2003)
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